FURTHER OBSERVATIONS ON C^SCAL PARASITES OF FOWLS. 75 
From the anterior blunt end four long free flagella take 
their origin. 
Running down one side of the body for a distance of about 
one third of the animal’s length from the anterior end there 
is a single row of darkly staining chromatic blocks, which 
seem in most cases to obscure a thin, short, chromatic line. 
This line is shown in PI. 13, figs. 52, 53. 
The blepharoplast appears to consist of four basal 
granules, each of which gives rise to one of the four free 
flagella. 
The nucleus is an oval, darkly staining mass lying near the 
anterior end. Usually the chromatin is scattered more or 
less irregularly through the substance of the nucleus. 
An axostyle is typically present, taking its origin from the 
blepharoplast, to terminate as a pointed rod at the animal’s 
posterior end. In addition to the elongate active forms, more 
rounded active forms are also met with in which the axostyle 
is curved round at its posterior end (PI. 13, fig. 55). In 
nearly all these forms, two small darkly staining wedges 
are found on either side of the axostyle near its posterior end. 
We have been able to get a fairly complete series of 
divisions of the active form of Trichomastix gallinarum, 
though here again we met with a curious time-variable in the 
behaviour of the various parts of the divided organism to 
which we have drawn attention above in the account of the 
division of the other forms. 
The division is here, as far as its initial stages are con- 
cerned, a longitudinal one. In the earliest stage of division 
that we figure (PI. 13, fig. 57) the blepharoplast complex has 
divided, two of the basal granules with their two attached 
flagella have remained in the old position near the anterior 
pole in connection with the old chromatic line. The other 
two, with their flagella, have moved to a new position in a 
slightly posterior oblique direction, and have given rise to a 
new chromatic line. The two new blepharoplasts are still con- 
nected by a chromatic band, which is probably analogous 
with a similar structure described in the division of Triclio- 
