276 
GUY C. ROBSON. 
carapace, after which the blood becomes pale. The animal 
in this condition has a bloated appearance, markedly different 
from the fresh pale condition of the stage succeeding a moult. 
I think we may conclude that some sort of fatty substance 
is lodged in tbe tissues in question, and assists in the for- 
mation of the new epidermis. The fate of the lipochrome can 
only be vaguely suggested ; probably it is got rid of at the 
moult, as the colourless condition of the new skin would 
suggest ; but I have never examined the freshly-cast carapaces 
to satisfy myself upon the point. 
The role of the yellow pigment is but generally indicated by 
the fact that the fat it accompanies is stored up in the ovary. 
It is certainly specific for this function, but its actual role, 
if it has any, is not necessarily indicated by the association. 
One is tempted, of course, to assign to each of these two 
lipochromes a differentiated physiological function and 
identity. Maly (4) found that they could be separated by 
appropriate chemical reactions from each other in the yolk of 
the ovary of Mai a squinado. But it is permissible to 
observe that while the blood of a mature female Inachus is 
more often yellow, the ripe ovary of the same animal is 
orange (pink -f yellow). Finally, when we take into con- 
sideration the passage from pink to yellow in the blood 
lipochrome (p. 275), it will be seen that, on the other hand, 
there is an equal possibility that these two colouring bodies 
are not rigidly determined each from each. 
IY. A Note on the Origin of the Fat Supply of Inachus 
in Relation to Sacculinisation. 
There seems to be little reason to doubt that the fat con- 
tained in the liver-cells of Decapoda is directly supplied by 
the food-stuffs of the animal. In addition to the experience 
of other autliors I may quote my own results on this point. 
After a forti light’s or eighteen days’ starvation fat begins 
to disappear from the liver. If a number of crabs which 
have been thus starved are taken and fed on a fat-con- 
