196 
J. BEARD. 
to be mentioned elsewhere as the ciliary ganglion. A figure 
such as Professor His gives in his recent work of two fibres 
passing out from two nuclei in the anterior cornu of the head, 
outside which they receive an “ Ehren-Wache ” of four 
“ parablastic ” cells, two on each side, is one which, in spite of 
much search, I have never seen. On the other hand, the 
figures I give in PI. XIX could be multiplied by the dozen, and 
figs. 53 and 58 are representations of an appearance which I have 
often met, and which Dr. van Wijhe assures me he also has 
very frequently seen. From these facts, and from facts regard- 
ing the development of anterior roots of cranial nerves, and 
nerves of the sense organs of the lateral line or branchial sense 
organs, I do not for a moment hesitate to declare that the 
facts of development are contradictory to (1) Professor His’s 
view of the absence of nuclei in the anterior roots, and (2) his 
assumption that when such nuclei are present they are of 
“ parablastic ” origin. It is worthy of notice that in his 
original assertions Professor His absolutely (vide supra) 
denied the presence of nuclei in the anterior roots, and only 
now that their presence in those structures in Elasmobranchii 
is obvious to every observer, including Professor His, does it 
occur to him to make use of that wonderful doctrine of para- 
blast to explain their presence. Professor His’s attempt to get 
out of a false position here is only a little more dignified than 
his endeavour to explain away the meaning he attached to the 
Zwischenstrang. 
IV. The Ganglionic Development in Different 
Grouts of Vertebrates. 
Without anticipating the results of my researches on other 
groups, which so far include Teleostei, Lizards, Frog, Newt, 
and Rabbit, I may be at least allowed to say now that the above 
mode of development of cranial and spinal ganglia holds, with 
very slight and unimportant modifications, for all these forms 
also. I might have left these forms undcscribed but that for the 
certainty that some observer or other would by-and-by quote 
