198 
J. BEARD. 
ledge is their doubt of the accuracy of Spencer’s and my in- 
vestigations on the origin of the root of a cranial nerve in the 
Frog. I have admitted the error above, and need not here 
mention the matter further. To one assumption of these two 
authors (No. 38, p. 11) I must, however, be allowed here to 
reply. They remark : “ More recently the theory of the de- 
rivation of the whole or greater part of the cranial nerves from 
the epiblast has been supported by Mr. Spencer and Mr. Beard. 
This view is a revival of that held by Gotte.” 
(1) The origin of a part of each of the cranial ganglia, and 
of what I called the suprabranchial nerves, was no longer a 
theory after the publication of my paper on the branchial sense 
organs (No. 6). It was then demonstrated for certain parts 
of the cranial ganglia 1 and for certain nerves that they have 
an epiblastic origin, and the matter could for these hardly be 
called a “ theory.” I can now demonstrate that the whole of 
the components of the various cranial ganglia are epiblastic in 
origin, and not wholly or in part outgrowths of the central 
nervous system. 
(2) Gotte never held this view, whatever may now be the 
case. I can only suppose that the two ladies never read the 
passages in his work which bear upon the question. The fol- 
lowing quotation from Gotte’s ‘Unke’ (No. 22, p. 719) gives 
a clear statement of Gotte’s conclusions at that time : — Bei der 
Untersuchung der Kopfnerven handelt es siclx zunachst urn 
ihre Zugehorigkeit zu den ganzen hintereinander liegenden 
segmentalen Abtheilungen des Kopfes ferner um ihre Unter- 
scheidung nach dem Ursprunge aus dem inueren oder aiisseren 
Segmente des mittleren Keimblattes oder aus andcren Era- 
bryonalanlagen jeder Abtheilung. Zu den letzteren gehbren 
der Sehnerv und die Seitennerven als Erzeugnisse des oberen 
Keimblattes, die ubrigen Kopfnerven eustchen aus dem 
mittleren Keimblatte.” 
1 I was inclined then to regard the whole of the ganglion as arising from 
the epiblastic sense thickening, and the cells derived from the “ neural crest ” 
as forming the root of the nerve. The point is a very difficult one to decide, 
and I refer the reader to a discussion of it in another part of these researches. 
