466 
DR. E. KLEIN. 
a superficial intraepithelial network^ the former situated in 
the layer next to the deepest columnar cells, the latter below 
the most superficial layer. The fibrils of the latter branch 
repeatedly, and run a horizontal course sometimes for con- 
siderable distances ; they join, cross each other, and wind 
their way between the epithelial cells, changing their level 
several times. 1 have also stated that the end-knobs that one 
meets with on some of these fibrils cannot really possess this 
character, since they are absent in others ; and, besides, they 
are identical with those larger varicosities that occur in the 
xjourse of most of the fibrils. 
Hoyer Archiv f. mikr. Anat.,^ Band ix, p. 234) describes 
the intraepithelial fibrils in the same way as I had done, 
but, while admitting anastomoses of these fibrils, main- 
tains that they terminate with free ends amongst the 
superficial layers of the epithelial cells. The small and large 
varicosities which these fibrils show in some specimens are 
absent in others, and, therefore, must be regarded as pro- 
duced artificially. Rollett (^Strieker’s Handbook,’ article 
Cornea,” p. 1136) does not see any anastomosis amongst the 
intraepithelial fibrils, but says, that having branched they 
terminate in free ends, often with a slight enlargement 
amongst the superficial epithelial layers. Waldeyer (Graefe 
and Saemisch, ^ Augenheilkunde,’ p. 210), on the other hand, 
maintains a terminal network of the intraepithelial nerves. 
Krause Allg. und Mikr. Anatomie,’ 1876, p. 539) saw the 
intraepithelial nerve fibrils forming plexuses near the surface; 
they terminate with small end-knobs. Izquierdo (1. c., p. 
27) finds that the intraepithelial nerves, having ascended in 
a vertical or oblique direction into the superficial layer, bend 
off into a horizontal course and branch. They do not 
anastomose. All fibrils terminate with free ends, or with a 
slight swelling. 
Waldeyer (^Archiv f. mikrosk. Anatom,’ Band xvii, p. 379), 
while accepting Izquierdo’s conclusions, admits the incorrect- 
ness of his former view of an intraepithelial terminal net- 
work. When speaking of a terminal network (Graefo and 
Saemisch, p. 270), Waldeyer then observed also free ends, but 
the possibility always remains that in such apparently free 
ends there is only an imperfect action of the chloride of 
gold.” This is just what I have urged in my first paper in 
this Journal, 1871, knowing well the great differences in 
the number of the intraepithelial nerve-fibrils that one meets 
in various specimens, and in various places of the same 
specimen, prepared after the same plan, and seeing that in 
