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with these fine nerve-fibres are oval masses of germinal 
matter or nuclei, which, with the fine nerve-fibres themselves, 
have been usually regarded as “ connective-tissue corpuscles.” 
2. That neither the epithelial cells of the frog’s tongue 
generally, nor those covering the simple papillae, are connected 
with nerve-fibres. 
3. That the mass consisting of epithelium-like cells upon 
the summit of the fungiform papilla is connected with the 
nerve-fibres, but it is not an epithelial structure. 
4. That the dark-bordered sensitive fibres constituting the 
bundle of nerves in the axis of the papilla divide near its 
summit into numerous very fine branches, with which nuclei 
are connected. Thus is formed a plexus or network of 
exceedingly fine fibres upon the summit of each papilla ; 
from this network numerous fine fibres may be traced into 
the special nervous organ, composed of epithelium-like cells 
upon the summit, with every one of which nerve-fibres appear 
to be connected. 
5. That the bundle of nerve-fibres distributed to a papilla 
always divides into two bundles which pursue opposite direc- 
tions. The division of the bundle may take place just at the 
base of the papilla, or at some distance from it, but it always 
occurs. 
6. That fine pale nerve-fibres pass from the same trunk of 
dark-bordered fibres as that which gives off the bundle of 
nerves to the papilla. The fine fibres ramify — 
a. Amongst the muscular fibres of the tongue. 
b. Upon the vessels. 
c. In the connective-tissue of the tongue generally, and 
also in the simple papillae. 
7. That the fine ‘ nucleated ’ nerve-fibres ramify freely 
amongst the delicate branching muscular fibres of the papillae, 
and form plexuses or networks which exhibit no nerve-ends 
or terminations, nor in any case does a nerve-fibre penetrate 
through the sarcolemma or investing tissue of the fibre, or 
connect itself with the nuclei of the muscle. As many of the 
muscular fibres are so very fine and narrow, the distribution 
of the nerves, and their exact relation to the contractile tissue, 
can be demonstrated very distinctly in the case of the muscles 
of the papillae of the frog’s tongue. 
