LOXOSOMA LOXALINA AND LOXOSOMA SALTANS. 129 
proximal diverhicula of long cells with small nuclei at their 
bases and filled with fine grannies of quite a different 
character to the cells of the larg'er distal pair. The cells of 
the apical part of the stomach and the intestine contain many 
granules at their basal ends which are blackened by osmic 
acid and are probably fat-globules. This part of the stomach 
may be i-egarded as absorptive (figs. 11 and 12). The 
characters of the proximal pair of diverticula suggest secre- 
tion of some digestive juice, the cells being filled with very 
fine granules not unlike the gi-anules of vertebrate pancreatic 
or salivary gland-cells. The margin of these cells or of some 
of them are irregular like those of mucous cells or other 
actively secreting cells, and probably indicate the pouring out 
of secretions into the general digestive cavity. These sur- 
faces are devoid of cilia. 
On the other hand, the larger, thicker-walled diverticula 
on the more ventral surfiice, which are yellow in the living 
animal, are formed of long large cells loaded with spherical 
masses of various kinds, some staining purple with haema- 
toxylin, others yellow and various shades of brown. Some 
stain intensely with thionin, others not at all. 'I’he char- 
acters indicate active constructive metabolism, and suggest a 
function to these parts of the alimentary tract more com- 
parable to that of a liver. 
The intestine has a narrow lumen, is circular in ti-ansverse 
section, and has a thick, homogeneous wall richly ciliated 
until it reaches the hood of the lophophore. Here the lumen 
widens, and fa3ces are collected, and the walls contain yellow 
pigments probably of an excretory nature. The anus is 
extremely small. I presume that it exists, but in no section 
have I been able to see it as an actual opening. Its position 
is indicated, however, by a [)roctodaeal depression and a 
suture which is probably caused by its coalesced edges. 
Nervous System. — The central nervous system, as in L. 
loxalina, is a conspicuous object lying between the intestine, 
and reproductive gland (fig. 14, n.y.). It consists of a pair of 
ganglia connected by a thick band of nerve-fibres. Peri- 
VOt. 58, PA({T 1. NEW SERIES. 9 
