200 
H. M. WOODCOCK. 
(27) and Schellack and Reiclienow (32) have maintained that 
no division of tlie karyosorne occurs in the above phase of 
13arrouxia (“Adelea”), and consider that the secondary 
karyosomes (i. e. the daugliter-karyosoines) arise de novo, by 
independent formation from the general nuclear substance ; in 
regard to this detail the authors are certainly mistaken. More- 
over, quite recently Chagas (loc. cit.) describes and figures, 
in his account of Chagasia (Adelea) hartmanni, per- 
fectly similar promitotic divisions of the karyosorne in 
different phases of the life-history. I have a strong idea that 
Reiclienow and Schellack, in arriving at the above conclusion, 
have been influenced — if unconsciously — by the view which 
one of them (Reiclienow) seems to have formed upon the 
question of the karyosorne, its nature and significance, as a 
result of his work on Haim ogre gar in a stepanovi. No 
one is more sensible than am I of the great value of 
Reichenow’s researcli, which has thrown full light upon the 
complicated subject of the Haemogregarine life-cycle ; but in 
regard to this somewhat important cytological question I find 
myself obliged to differ from him. 
Hartmann and Chagas (10) have suggested that the reason 
for this may be that as the particular parasite (H aem ogre- 
gar ina stepanovi) upon which Reiclienow worked is a very 
small one, tlie observation of minute cytological details and 
clianges would be rendered more difficult and hence they may 
have escaped detection. I do not altogether share this opinion ; 
for one thing, I do not think H. stepanovi is much, if any, 
smaller than the small forms of K. lacertae, where thekaryo- 
some and its division can be made out without difficulty. I am 
inclined to consider that, on the whole, the nuclear constitu- 
tion and behaviour in H. stepanovi is as Reiclienow has 
described it ; and therefore, as a logical sequel, that this 
species of Hamiogregariiie differs in one or two cytological 
respects, such as the absence of a typical karyosorne, from 
certain other Hmmogregarines and certain Coccidia. This is 
the more probable, in my opinion, because of a fact which is 
evident on scrutinising Reichenovv’s figures, namely, that the 
