MUSCLES INNERVATED BY THE FIFTH CRANIAL NERVE. 601 
branches, tliongh examination by the Marchi method has 
failed to reveal them.” 
Willems (’ll) also found chromolytic changes in the mesen- 
cephalic nucleus after avulsion of the individual motor 
branches of tlie fifth. 
Though the observations described in this paper show the 
existence of ganglionated afferent nerve-fibres in the muscle- 
branches of the fifth nerve, they leave untouched the difficult 
question of the peripheral distribution and function of the 
axons of the mesencephalic root. 
I owe many thanks to Sir Victor Horsley for performing 
the operations described above. The expenses have been 
defrayed by a grant from the Bristol University Colston 
Committee. 
September 18th, 1912. 
Referexces. 
Beevor and Horsley (’88) — “ Note on some of the Motor Functions 
of certain Cranial Nerves (fifth, seventh, ninth, tenth, eleventh, 
twelftli), and of the first three Spinal Nerves in the Mtinkey 
(Macacns sinicus),” ‘Proc. Roy. Soc.,’ vol. xliv. 
Cipollone (’97) — “ Richerche suH’ anat. d. terminazone nervose nei 
mnscoli striati,” Roma. Cited by Shenington, in ‘ Text-book of 
Physiology,’ ed. Schafer, vol. ii, 1900. 
Cords, Dr. Elizabeth (’10) — “ Ziir Morphologie des Gaumensegels,” 
‘ Anat. Anzeig.,’ September 3rd. 
Davies (’07) — “The Functions of the Trigeminal Nerve,” ‘Brain,’ 
vol. XXX. 
Dogiel (’00) — “ Die Endingungen der sensiblen Nerven in den 
Angenmuskeln nnd deren Sehnen beim Menschen imd den Siiu- 
gethieren,” ‘ Archiv f. microscop. Anatomie,’ vol. Ixviii. 
May and Horsley (’10) — “ The Mesencephalic Root of the Fifth 
Nerve,” * Brain,’ part cxxx. 
Sappey (’72) — ‘ Traite d’ Anatomie descriptive,’ vol. iii. 
Shen-ington, ('91— ’95) — “ On the Anatomical Constitution of the Nerves 
of Skeletal Muscles ; \\-ith Remarks on Recmaent Fibres in the 
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