STUDIES IN THE EXPERIMENTAL ANALYSIS OF SEX. 583 
all Grossu lariata. This is the generation used in 
succeeding crosses. 
(2) F^ grossnlariata ? x Fj gross u lariata S gave 
Fj grossular iata s and $ s and lacticolor ? s, but no 
lacticolor s. 
(3) Lacticolor ? x F;^ grossulariata ^ gave grossu- 
lariata d* s and ? s and lacticolor cJsand ? s, the lacti- 
color J s being the first ever seen. 
The explanation of these results offered by Bateson and 
Punnett depends on the two assumptions — 
(1) Tliat the female is heterozygous in sex, femaleness 
being dominant, and the male a homozygous I’ecessive. 
(2) That when in Fj the two dominant characters, female- 
ness and the grossulariata factor co-exist, there is spurious 
allelomorphism or repulsion between them, such that each 
gamete takes one or other of these factors, not both. 
The three crosses described above read, therefore : 
(1) Lac. ? X (fros. S 
Produce gametes — 
Lac. ? . Lac. x Gros. S 
Produce zygotes — 
Fi Lac. ? Gros. S • Gros. S • 
(2) Fi Gros. ? X Fj Gros. S 
Produce gametes — 
Lac. ?. Gros. J (Lac. d' ) (Gros. ?) x Lac. J (iros. S 
Produce zygotes — 
Lac. ? Lac. cJ . Lac. ? Gros. J . Gros. d* J . Gros. c? 
Gros. J . 
The gametes in brackets are not formed owing to spurious 
allelomorphism. 
(3) Lac. ? X Fj Gros. J 
Produce <>:amete.s — 
Lac. ? Lac. ^ x Gros. c? Lac. c? 
Produce zygotes — 
Lac. ? Gros. J . Lac. ? Lac. . Lac. J Gros. . 
Lac. cJ Lac. cJ . 
This remarkable case has been given in full, as it illustrates 
