34 
THE VOYAGE OF H.M.S. CHALLENGER. 
close to those organs “ die seit Cuvier fiir die Speicheldriisen gelten.” For the rest he 
does not say what is his own opinion in regard to the nature of these bodies. 
I do not think that since the publication of Darwin’s Monograph these organs have 
been investigated ; so I was most anxious to study them, and if possible to make out 
their structure. They occurred in all the genera in which I sought for them ; I studied 
them in greatest detail in the genera Lepas and Scalpellum. 
Near the place where the oesophagus communicates with the stomach, the outer surface 
of this latter organ is invested with a pair of oval masses ; they are placed at rather a 
considerable distance from one another, one being found at the right, the other at the 
left hand side of the stomach. PI. VI. fig. 7 shows their situation in Lepas anatifera when 
seen laterally, fig. 8 when seen from the anterior (dorsal) side. In both figures CE . repre- 
sents the oesophagus and G. S. the supracesophageal ganglion ; p. n. are the two strong 
peduncular nerves which start from the supracesophageal ganglion ; oc. is the curious eye 
discovered by Leidy, placed close to the surface of the stomach and separated from the 
external surface of the body by a very darkly pigmented integument and a thick layer of 
muscles, which are both left out in the figures. The oviducts ( ov ) are also distinct in both 
figures. They come from the peduncles and for some distance run parallel to the pedun- 
cular nerves ; a little beyond the eye they are seen to diverge and then may be followed 
running transversely over or at least close to the surface of the stomach. Dorsally from the 
oviducts (in fig. 7 beneath them) the most anterior parts of the testis (t) can be distin- 
guished. That part of the surface of the stomach which is nearest to the oesophagus is 
covered all over with rounded and dark-coloured tubercles ( l ) which cause the “ disposition 
mamelonnee” of Martin-Saint- Ange, and which when studied in a transverse section 
appear to be the arborescent coeca of the surface of the stomach. The internal surface of 
these coeca is darkly pigmented, and this causes the blackish colour of the rounded 
swellings at the exterior. 
The glandular bodies in figs. 7 and 8 are marked gl. They are not always of the 
same shape and size. Sometimes they are rather regularly oval and compact, having a 
length of about 4 mm. and a breadth of not quite 2 mm. In other cases, however, finger- 
shaped excrescences (as observed by Darwin) give the gland a much more irregular 
appearance. In both cases the surface of the body is uneven owing to the presence 
of globular swellings ; whilst the whole body represents an acinous gland, each of the 
globules being a distinct acinus. 
Before giving a description of the microscopic structure of the gland in Lepas I will 
describe its structure in Scalpellum. My best preparations are from Scalpellum parol - 
Idogramma , Hoek. In this species the gland is relatively small, having a length of little 
more than one millimetre. It is pyriform ; at the narrow extremity it communicates 
with the interior of the stomach by means of a very narrow duct ; at the other extremity 
its body is rather blunt and rounded. The greatest transverse diameter of the gland in 
