— 7 i— 
ing below the point and on the very narrow base of the dorsal lamella. Vag- 
inant lamina rather more than halE the length of the leaf-complex. Cells of 
middle and upper part roundish hexagonal to polygonal, 6-11/4 in diameter, 
often quite variable: basal celis rather hyaline, quadrangular, some nearly 
square, 13-27 X 6-10/*. Synoicous or dioicous, the flowers terminal or some- 
times in a bud near the base. Antheridia 1-5 (1-7) large, subclavate, the 
slender archegonia more numerous, sometimes 18-20, without paraphyses. 
Seta reddish, 3-4 mm. long. Capsule erect, green, oval-oblong to subcylin- 
drical, o. 5-0.7 mm. long, the mouth purplish. Operculum conic-rostrate, 
long-beaked, the length of the capsule, straight or slightly oblique, 
brownish. Teeth russet-brown, broad, rather short, not deeply divided 
smooth or little papillose, the broad cross-bars horizontal. Spores globu- 
lar, green, pitted, 13-16/4 in diameter. 
Moist and shaded calcareous rocks. Fraction Run (now Dellwood Park) 
Lockport, 111 ., June, 1905, E. J. Hill; San Marcos, Texas, Charles Wright, 
1847. At Lockport it is associated with Gymnostomum calcareum Br. Germ 
and A mblystegium varium (Hedw.) Lindb. 
The chief difference between F. inconstans and F. synoicus is in the 
inflorescence and the time of maturing the spores, winter or early spring in 
one, early, or midsummer in the other. This would of itself suffice to' differ- 
entiate them specifically. The teeth of the former are slender and quite 
long, divided below the middle, the parts very papillose and spirally thick- 
ened. In F. synoicus the teeth are not deeply divided, smooth or smoothish 
the parts not spirally thickened. In F. inconstans the thickened leaf-border 
is generally confluent with the costa at the point and readily separates from 
the cells of the body of the leaf; in F synoicus the border is less developed 
or more interrupted, not thickened or cord-like, and usually ceases below the 
point. Its spores are green and pitted, those of F. inconstans are yellow or 
brownish yellow. 
In its inflorescence F synoicus is nearest to F. Bambergeri. Schimper 
described this as dioicous, (Syn. Ed. 11, p. 115). Roth, Boulay, Husnot and 
Milde give it variously as polygamous, synoicous, dioicous. I have found it 
both synoicous and dioicous. Of twelve stems examined eight had floral 
organs, two of them synoicous, two had archegonia only, four antheridia or 
with buds having the short leaves of the perigonial form. In synoicous flow- 
ers the leaves were the larger, perichaetial form, with 2-5 archegonia and 
one or more but few large antheridia, all kinds without paraphyses, or rarely 
with some. In the case of stems without fruit my experience is similar to 
that of Milde, who says: “I always found the fruiting plants without male 
flowers. The plant is also dioicous. The sterile plants bear either a single 
female terminal flower, which consists of 5-12 archegonia without para- 
physes, or a single terminal hermaphrodite flower in which I found r-3 
antheridia and 3-5 archegonia without paraphyses. Since the antheridia 
were always very large an error is not possible. The archegonia on the other 
hand were always very long and slender.” (Zur Kryp. Flora sud. Tirol. Bot 
Zeit. 22 : 12, 1864.) Milde considers it similar to F. incurvus : Schimper as 
