274 
Jenssen 
pauses which determine the bob cadence and 
serve as the major pattern-producing factor. 
As a case in point, the data from 33 filmed 
displays of a typical male opalinus show the 
displays varying from five to eight bobs 
(Fig. 4), with a range of head-bob durations 
of 1.7 to 4.3 s, and a C.V. of 23.3 percent. 
Degree of Display Complexity 
Anolis lineatopus neckeri from luandeville, 
Jamaica, has a very short and stereotyped 
signature display (Fig. 5). The pattern, 
derived from 93 filmed displays by four 
males, consists of a short series of continu- 
ous sine wave bobs (Jenssen, 1977a). The 
total duration of the display is just over one 
second. 
In contrast to A. 1. neckeri, the signature 
display of Anolis sericeus from Vera Cruz, 
Mexico, is usually long, averaging almost 
30 s, and is quite complex. Closely coordin- 
ated dewlap pulses are integrated with a 
complex head bobbing pattern (Fig. 6). 
Based on 21 filmed displays of five males, 
the pattern is quite predictable except for 
some introductory head movements and a 
terminal dewlap pulse (nondarkened areas 
in Fig. 6). This display is about 30 times 
the duration of the signature display of A. 1. 
Figure 4. Distribution of bobs within displays hav- 
ing the same number of bobs/display (6-bob to 
8-bob displays) by a male Anolis opalinus. Head- 
bob amplitude is stylized, with the interbob pauses 
expressed as a percentage of the total display 
duration. The thin horizontal lines denote one 
standard deviation on either side of the mean 
inter-bob pause values (medial vertical lines). 
neckeri. The greater variety of muscular 
movements and their nonrhythmic coordina- 
tion are in marked contrast to A. 1. neckeri' s 
cyclic and continuous sine wave bobs. 
Size of Display Repertoire 
In analyzing displays, I treat a species’ 
repertoire size conservatively, restricting the 
count of different display types to only those 
body movement patterns which are stereo- 
typed and species-unique (Jenssen, 19776). 
According to such criteria, Anolis nehu~ 
losus from Nayarit, Mexico, has only one 
display type (Fig. 7a). I originally called this 
stereotyped pattern the “assertion” display 
(1970; 1971) but, keeping with more recent 
terminology, it is more appropriately labeled 
as the species’ signature display. 
r I 1 — 
0 I 
SECONDS 
Figure 5. DAP graph of the signature display of 
Anolis lineatopus neckeri. Upper edges of the 
blackened area depicts head amplitude movement 
through time. Stippled area represents a different 
bob pattern sometimes associated with the signa- 
ture display. 
▼ ▼ ▼ 
5 10 15 20 25 30 
SECONDS 
Figure 6. DAP graph of the signature display of 
Anolis sericeus. Upper block depicts head ampli- 
tude movement, and lower block represents dewlap 
extension. Blackened areas are stereotyped por- 
tions of the display, and white areas are examples 
of variable portions of the pattern. 
