Table 56. — Social rank and important conditions associated with it 
Rank 1 
Number 
Mean May 
1949 wt. gr. 
Mean life-span 
maturity index 
Mean number 
of wounds 
Placental scars 
Young 2 
Number 
Mean 
Males 
Females 
Males 
Females 
Males 
Females 
Males 
Females 
of females 
examined 
per 
female 
Total 
Average 
female 
High 
7 
24 
510 
464 
1.73 
1.63 
10.4 
5.2 
15 
13. 1 
95 
3. 96 
Medium 
12 
17 
479 
431 
1.92 
1.96 
11.4 
5. 5 
9 
12. 9 
37 
2. 18 
Low 
42 
18 
441 
419 
11.09 
11.19 
15.4 
6.4 
13 
5.4 
0 
0. 00 
1 See figure 128 for the grouping of the colonies according to high, medium, or low rank. 
2 Bom March to May 1949, and still alive May 17, 1949. 
Table 57. — Indices of stress 
Colony 
1 
Weight 
index 
2 
Proportion 
females 
3 
Homogeneity 
index 
4 
Proportion 
of females 
reproducing 1 
Average num- 
ber of young 
per female 
5 
Index of 
production 
of young 
Composite 
index (mean 
of 1 to 5) 
a 
0. 884 
0. 928 
0. 90 
0. 769 
4.3 
1. 000 
0. 896 
b 
.698 
.666 
.91 
.750 
3. 8 
.883 
.781 
c+d 
.776 
.600 
.75 
.428 
3.4 
.790 
. 669 
e 
.648 
.642 
.72 
.555 
2.3 
. 537 
. 620 
f 
.378 
.534 
.69 
. 250 
2.0 
.465 
.463 
S 
.228 
.812 
.65 
. 153 
0.0 
. 000 
. 369 
h 
.202 
.500 
. 81 
.000 
0.0 
.000 
.302 
.056 
.000 
.71 
. 000 
0.0 
.000 
. 153 
j 
. 294 
. 100 
. 47 
2 1. 000 
0.0 
.000 
. 373 
k 
.000 
.000 
.53 
.000 
0.0 
.000 
. 106 
1 Lactating or pregnant May 1949. 
3 Only 1 female in colony. Though she was lactating when examined in May 1 949 she had already lost her recently born 
young. 
low homogeneity of origin approaches zero. Low 
homogeneity indices reflect forced dispersal, and 
lack of stable social oranization. A wider variation 
in this index would arise were it feasible to 
incorporate into it data concerning proximity to 
other colonies of known social rank. 
4. Proportion of females reproducing. 
5. Index of production of young. 
The most successful colony reared on the average 
only 4.3 young per female. An index of 1.0 was 
assigned to this colony. The average number of 
young for the others are represented as percentages 
of 4.3. An arbitrary index of zero was given to 
those colonies lacking females since all indications 
were that even had females been present no young 
would have survived. 
In the rank ordering of the colonies from a to k 
in a descending order of social rank, it may be seen 
that, with the exception of colony j, at least two 
out of the three indices are lower than for the pre- 
ceding colony (table 57). No single one of these 
indices is alone adequate for judging the relative 
amount of stress. As the amount of stress increased, 
there was a decline in the reproductive perform- 
ance of females. This decline is reflected both in 
the proportion of the females who were lactating 
or pregnant, and in the average number of young 
per female. 
Reproduction is inhibited in two ways (see table 
56). First, as the social instability and the result- 
ant stress begins to increase, there is a reduced 
ability to rear the young which are bom. This 
215 
