258 
J. STEPHENSON. 
the author could not find that they were in any way connected 
with the pharyngeal system of glands. 
The pharyngeal and septal glands ofAleodrilus keyesi 
are also described. Here it will be sufficient to call attention 
to the author’s statements regarding the discharge of the 
gland-cells. The pharyngeal glands have discharge pockets 
which are much thicker than those seen in any other species; 
the septal glands are of the same nature as the pharyngeal, 
“ but I have good reasons to believe that the glands in this 
species discharge into the tubular intestine. I have been 
able to follow fclie discharge duct as far as the muscular 
layers of the intestine, which would hardly have been the 
case if the ducts had continued forwards into the pharynx, 
as do those of the forward septal glands in many genera.” 
In some other small aggregations of similar cells the author 
was unable to follow the ducts. 
In Sparganophilus (which, though aquatic, belongs to 
the Grlossoscolecidse, and so may be considered along with 
the earthworms), it is noted that in one species the ducts of 
the septal glands with precipitated secretions can be followed 
along the septum down towards the intestine, but the con- 
nection with the latter, if any, was not ascertained; in 
another species the discharge tubes and chambers are very 
large, the chambers occupying more than half the width of 
the pharyngeal wall (the meaning is more than half the 
height of the pharyngeal epithelium). 
De Ribaucourt (8, 1901), describes in a few words the 
deeply staining mass of cells in the Lumbricidas : “ On 
staining with methyl blue and iodine green one can easily 
establish the fact that these cells are continued as far as the 
epithelial layer by a fine prolongation ; thus the cells may 
quite possibly have a secretory function.” Miss Raff (7a, 
1910), recognises the cells in the Australian Megascolecidae, 
but finds no trace of a duct in connection with the “ glandular 
mass.” 
I omit the literature which deals with the septal glands of 
the specially aquatic groups — the Microdrili — as 1 hope to 
