PHARYNGEAL GLAND-CELLS OF EARTHWORMS. 
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example, Megascolides, Notoscolex, Megascolex, 
Lampito, Pheretima, Ery thrseodrilus, Octochaetus, 
Eutyphoeus, Eudi chogaster — to mention only genera in 
which I have myself observed them. The nephridia of 
meganephric forms are not enlarged in the pharyngeal and 
immediately subsequent segments ; why no modification of 
any kind occurs in them, when in other and sometimes closely 
related genera a great multiplication and massing together of 
the micronephridial tubules takes place in this region, I am 
unable to say. 
The chromophil cells do not stain with Lugol’s iodine 
solution; glycogen seems therefore to be absent. 
Summary. 
(1) The “ pharyngeal gland-cells ” of earthworms are not 
gland-cells in the usual sense, and do not communicate with 
the pharynx ; the term “ chromophil cells ” is proposed for 
them because of their intense coloration by haematoxylin 
and similar stains. The so-called “ septal glands ” of earth- 
worms are aggregations of similar cells at a more posterior 
level. 
(2) In the chromophil cells the deeply staining matter is 
not equably distributed through the cell-body ; the peripheral 
regions of the cells in general stain more lightly, and appear 
to be disintegrating, or merge into an intercellular substance. 
(3) While most of the cells form a more or less compact 
aggregate on the surface of the pharyngeal mass, a number 
penetrate inwards towards the pharyngeal epithelium, and 
become progressively metamorphosed into fibrillar connective 
tissue. 
(4) A capsule of flattened cells covering the mass, though 
present in part, is incomplete. The smaller masses of cells in 
Helodrilus parvus are frequently continuous with the 
peritoneal membrane, of which they appear as modifications. 
(5) In Helodrilus parvus, and especially in all young 
earthworms, the inwandering and the connective tissue change 
