350 
EDWARD PHELPS ALLIS. 
the trapezius thus apparently representing the fifth levator. 
In Menidia, Herrick (1899, p. 117) found a trapezius inner- 
vated by a branch of the vagus, and there are but four 
levatores in that fish. In Scomber I found (Allis, 1903, 
p. 207) five external levatores, the fifth one being inserted in 
a membrane attached to the clavicle, and there is no trapezius 
in this fish. In Trigla and Scorpaena I also found (Allis, 
1909) five external levatores, and I now find that there is no 
trapezius in either of these fishes. In Ameiurus there is a 
trapezius innervated by a branch of the vagus (Herrick, 
1901, p. 209), and the levatores of this fish are none of th m 
inserted on the fifth branchial arch (McMurrich, 1884). In 
Polypterus, Edgeworth (loc.cit., p. 241) finds a trapezius, 
and there are but four levatores in this fish. 
These several facts regarding these fishes, when compared 
with Greil’s descriptions of Ceratodus, seem certainly to 
warrant the conclusion that in the Teleostomi, as in Ceratodus, 
the trapezius is developed from the fifth branchial myotome, 
and that it always retains its normal and primitive innervation 
by the nerve of that segment of the body. 
The musculi coracobranchiales are said by Dohrn to be 
developed, as already fully explained, from the deeper, 
proximal fibres of the ventral portions of the constrictores 
superficiales of the branchial arches and to be represented, in 
the adult, by those fibres as described by Vetter. They are 
accordingly said by Dohrn to be totally different muscle-s 
from the coracobranchiales of Vetter’s descriptions of the 
adult, which are said by Dohrn to be simply the distal fibres 
of the ventral portions of the branchial constrictores super- 
ficiales, misnamed coracobranchiales by Vetter. Edgeworth 
says, as already explained, that the coracobranchiales are 
developed from the entire ventral ends of the branchial 
myotonies, and he considers the muscles so developed to be 
identical with the coracobranchiales of Vetter’s descriptions 
of the adult. 
'Phe coracobranchialis of the first branchial arch of the 
