CYTOPLASMIC INCLUSIONS OF THU GEltM-CELLS. 455 
Meves does not describe the acroblasts, for he has not found 
them, but it would be of great interest to know what happens 
to these structures. 1 We know that the macromitosome is 
apparently normal in the “ apyrefpe ” sperm. This shows 
that the nucleus is not concerned in the formation of this 
cell organ ; probably the macromitosome, if I may use the 
expression, is “ looked after ” by the centrosome or centro- 
somes. The acroblasts, by their behaviour in growth and 
division, seem to rely on the nucleus ; we thus get the interest- 
ing state of different cell elements being independent inter 
se and governed by special parts of the cell. The centro- 
some governs the macromitosome, the nucleus influences the 
acroblasts, and the centrosome keeps close to the nucleus ; 
and so the elements are orientated correctly in the sperm. 
Exactly what part each plays it is quite impossible to state, 
and the extent in which cytoplasmic movements affect the 
question cannot be decided at present. 
The moth or butterfly never lacks sufficient normal sperma- 
tozoa, and it is not until a great number have been formed 
that the “apyrene” spermatozoa become formed in any 
number; but in many species in later stages the abnormal 
spermatozoa are exclusively found in process of formation. 
Von N. Divaz, in a late paper ( 14 ) on the “ Spermato- 
genese von Naucoris cimicoides,” treats of spermato- 
genesis from the spermatocyte at the end of growth stage 
onwards. He identifies, in the cytoplasm of the first 
spermatocyte, some three bodies, which he calls Archoplasmaj 
and several chromophile bodies (Chromatophile Korperchen). 
Since he has partially destroyed the mitochondria with acetic 
fixation, his account of the formation of the macromitosome 
is inadequate. In PL 1, fig. 1, he depicts, but does not 
mention, the mitochondria, which are in a dissolved con- 
dition. His archoplasmic corpuscles are probably acroblasts, 
and he figures them with a crescentic edge turned towards 
the nucleus, in the opposite way to that in which I found 
them in Lepidoptera. In the spermatid he believes that the 
1 See my paper on the “ Apyrene spermatozoa.” 
