Contributions to the Histology of Brady pus tridactylus. 39 
The complex stomach is divided into three groups of compart- 
ments, and the histological characters show that some have a 
secretory function, but the others are purely mechanical. The 
compartments are : — 
1. The paunch (A) with a conical csecal appendix. 
2. The cardiac stomach derived from the oesophagus and 
divided into three compartments (B, C, D). 
3. The U-shaped pylorus consisting of two tubular parts (E,F). 
Two of the compartments of the cardiac stomach are separated 
by a septum (G) which runs into the paunch for a short distance. 
Histology . — When a section is made through the septum (G) 
Text-Fig. 2. — The Stomach. I., divisions; II., view of a fold of 
the mucosa ; III., longitudinal folds running between glandular 
masses in the first part of the pylorus; R., duodenum. Other 
letters in text. 
at the point where it passes into the paunch, one sees the 
stratified epithelium of the cardiac stomach ( a ) giving way to the 
glandular epithelium of the paunch (b) (text-fig. 3) ; and the 
change resembles that of the gastro-cesophageal junction of a 
simple stomach. The transition is a sharp one, but does not take 
place at the same level on each side of the septum. The corium 
contains lax connective tissue, capacious blood-vessels (c), and 
aggregations of lymphoid cells ( d ). The muscular coats are 
diminished. 
The mucosa of the paunch (text-fig. 4) is succulent, pitted by 
innumerable glandular orifices and thrown into folds. A section 
? The Stomach (text-fig. 2) 
