444 
H. M. WOODCOCK AND G. IWPAGE. 
yefc proceeded along tlie entire lengthy and the two daughter- 
flagella are still united into one distally. Probably the 
parent-flagellnm splits nearly throughout its length, for the 
two daughter-flagella are usually approximately equal (figs. 
24-28) ; it is exceptional to find a dividing individual in 
Avhich there is as great a difference between the length of the 
separated flagella as in that of fig. 31. We have been consider- 
ably exercised in regard to the question whether there is a basal 
granule in connection with the flagellum. We are rather 
inclined to think that there may be such a granule, but we 
cannot say with certainty. There is frequently a definite 
granule exactly at the point where the flagellum originates 
(tigs. 11, 13, 21 and 22), but owing to the peripheral situation 
of the chromatinic zone, it is possible, of course, that such 
granule is a chromatin grain. Nevertheless, in such a case 
as is shown in fig. 20, where the chromatinic zone is very 
feebly developed, but where there is a very distinct granule- 
like thickening at the basal end of each of the daughter- 
flagella, definite basal granules are certainly suggested. 
As regards the division of the chromatinic substance, we 
have found nothing to indicate that there is any pronounced 
attempt at equal distribution between the two daughter- 
individuals. Apparentl}^, the chromatic substance which 
happens to be in either half, prior to division, goes to that 
daughter-individual (cf. figs. 24 and 25). However, the 
most usual condition in which the chromatin occurs in 
dividing individuals is that of a number of small granules 
fairly uniformly distributed around the periphery (figs. 27, 28), 
and therefore there may be really a nearer approach to 
equalisation than is obvious owing to the absence of a definite 
nucleus. The last act in the process is the constriction of the 
general body- substance into two halves ; this always takes 
place exactly in the middle of the long axis. 
In some cases fission of the body undoubtedly occurs before 
the flagellum has split (figs. 32, 30, 29 show different stages 
in such a process). Whether the daugditer-individual which 
thus lacks a flagellum is able to develop one we cannot say. 
