(ENOTIIERA MUr. LATA AND (L. MUT. SE.MILATA. 
527 
Solauder (Davis^ 1909), as well as in Ion gi flora (Beer, 
1906), Q^]. muricata, CK. c rue i at a and ffi. Miller si 
(Stomps, 1912). It will, therefore, be obvious from these 
facts, together with the description of the cultures above 
enumerated, that plants having the foliage characters of lata 
or semilata and 15 chromosomes arise sporadically from 
races having 14 chromosomes. 
CE. MUT. LATA AND SeMILATA. 
Before proceeding with the description of the plants in 
these cultures, it will be necessary to point out the chief 
peculiarities of lata and semilata. These forms were first 
described by de Yries (see 1909, vol. i, p. 415), in whose 
cultures lata appeared as a mutant from CE. Lamarckiana 
and several of its derivatives to the number of 493 plants in 
a total of 130,000 seedlings, or with a frequency of about 0’4 
per cent. The most reliable percentages obtained in families 
of 8000, 10,000 and 14,000 plants respectively were 0‘5 per 
cent., 1‘7 per cent, and T8 per cent., while in one small culture 
of 164 plants the number of latas ran up to 5 per cent. 
These fluctuating frequencies are doubtless an indication of 
environmental influence. 
In de Vrieses cultures, semilata only appeared in the off- 
spring of lata X Lamarckiana. OE.lata was classed by 
him as an inconstant species, but semilata was incorrectly 
classed as constant. They are both obviously inconstant, how- 
ever, and the presence of the odd chromosome shows why 
this must be so. 
Text-flg. 1 shows a typical rosette of lata, as it occurs in 
cultures from de Vries’s race of Lamarckiana. The leaves 
are broad, with usually very obtuse, rounded points, and very 
deeply crinkled. Lata also has a peculiarly hand-shaped 
stigma, the buds are stout and barrel-shaped, usually more 
or less symmetrical, with the petals crumpled inside the 
bud. The anthers are almost wholly sterile, and in typical 
plants there is a complete absence of red pigment from all 
