546 
It. RUGGLES GATES AND NESTA THOMAS. 
CE. biennis (1910). Tlie corresponding 15-13 distribution 
was found in CE. mut. gigas (Gates, 1911a) and a 9-11 
distribution in a 20-cliromosome plant, having lata as the 
mother (Gates, 1910). 
In fig. 37 a somewhat different condition from any observed 
in lata rubricalyx was discovered, namely, a tendency for 
the chromosomes to leave portions of their viscous substance 
trailing behind as they pass towards the poles. This condition 
was not infrequently observed in biennis lata. Itapparently 
denotes a pathological condition of the chromosomes con- 
cerned, and is probably followed by their degeneration. 
Geerts (1911) has described the fragmentation and degene- 
ration of certain chromosomes in the homotypic division in 
the pollen mother-cells of gigas x Lamarckiana, and in 
the first meiotic division of the megaspores of lata x 
2 'ioras. He also found that during interkinesis in tlie 
pollen mother-cells of the last cross, some of the chromo- 
somes (which would probably afterwards degenerate) failed 
to undergo the usual split. His figures also indicate that 
certain of the chromosomes may have split on the hetero- 
typic spindle as here described. 
L am arcki ana-1 ike Plant (142. 1.2). 
Figs. 42 to 45 show certain features of the meiotic chromo- 
somes in the Lamarcki ana-like plant (142. I. 2), having 14 
chromosomes, derived from self-pollination of semilata. 
The heterotypic spindle drawn in fig. 42 contains 14 chromo- 
somes, two of which are leaviiig behind trails of chromatin 
similar to those observed in biennis lata (fig. 37). Fig. 43 
shows the normal homotypic metaphase ’with 7 chromosomes 
in each group, while in fig. 44 an irregular distribution has 
taken place, the result of which is not quite clear. But there 
appear to be about 6 whole chromosomes and a fragment in 
one group ; and the other group contains 8 bodies whose 
valency is not clear in every case, though there are obviously 
great differences in the size of the chromosomes. In this 
