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TORTULA PAQORUM (MILDE.) DE NOT. IN GEORGIA. 
A. J. Grout. 
In working over my collection of Tortulaceae as a preparation for writ- 
ing up the family for “Mosses with Hand-Lens and Microscope ” I found 
that Dr. John K. Smalls’ “ Mosses of the Southern United States,” No. 39, 
on Ulmus, Atlanta, Georgia, July 1, 1895, labelled Barbula papillosa 
(Muell.) Wils. was not that species at all, and it was evidently no species 
credited to North America. 
The plants are short, 3-5 mm., stout, very dark or brownish green leaves 
when dry appressed and somewhat contorted, when moist erect-spreading, 
lingulate to obovate or fiddle-shaped, margins plane below entire, occasionally 
incurved at the middle, sometimes emarginate according to Limpricht, but 
in our specimens not emarginate and with the upper margins incurved as in 
T. papillosa as is shown in the illustrations, Plate VIII. Leaves strongly 
papillose above with papillae of various shapes and degrees of 
complexity as shown in the illustration; lower leaf cells much larger, rectan- 
gular, nearly hyaline; costa excurrerit, smooth at back. In the axils of the 
upper leaves are numerous leaf-like, ecostate broodbodies, having strongly 
papillose cells and a hyaline apiculus. 
Easily known from T. papillosa which has simple papilla on the leaf 
cells, larger forked papillae on the back of the costa and numerous 
clavate to subglobose, multicellular broodbodies clustered on the upper sur- 
face of the costa. The illustrations will serve to make the distinctions 
clear. T. papillosa is probably common on the bark of trees, and I expect 
that it will be frequently reported now that its determination is made easy. 
T. pagorwn is likely to be found much farther north and herbarium speci- 
mens labelled T. papillosa should be carefully examined for it. Both 
species rely on their broodbodies for reproduction. The fruit of T.pagorzan 
has never been collected, while the only fruit of T. papillosa known to 
science came from Australia and New Zealand. 
Plate VIII, Fig. 1. Tor tula papillosa, 1, Leaf showing brood bodies on 
costa, la, apex of leaf showing papillae and brood bodies. After Dixon and 
Jameson. 
Fig. 2. Cross sections of leaf showing brood bodies and papillae. After 
Limpricht. 
Fig. 3. Tortula pagorum, brood bodies highly magnified, “c”, cross 
section of costa much less magnified. After Limpricht. 
Fig. 4. T. pagormn , apex of leaf and marginal papillae. 
Fig. 5. Leaves and cells of same. Figs. 4 and 5 after drawings by Miss 
Thayer. 
