Date and Place of Birth: 
Arthur Wesley Nienhuis 
August 9, 1941; Hudsonville, Michigan 
Fducation : 
May 
January 
June 
June 
June 
June 
June 
1959 Graduated, Western Christian High School, Hull, Iowa 
1961 Cornell College, Mt. Vernon, Iowa 
1962 Los Angeles City College 
1963* College of Letters and Sciences, University of California, 
Los Angeles 
1965 Medical School, University of California, Los Angeles 
1966 Graduate School, Department of Anatomy, University of 
California, Los Angeles 
1968 M.D., with highest honors. School of Medicine, University of 
California, Los Angeles 
Brief Chronology of Employment : 
1965-1966 
1968- 1969 
1969- 1970 
1970- 1972 
1972- 1973 
1973- 1977 
1976- present 
1977- present 
Predoctoral Fellow, Department of Pathology, University of 
California, Los Angeles 
Intern, Massachusetts General Hospital, Boston, 
Massachusetts (Straight Medicine) 
Assistant Resident in Medicine, Massachusetts General 
Hospital, Boston, Massachusetts 
Clinical Associate, National Heart and Lung Institute, 
National Institutes of Health, Bethesda, Maryland 
Clinical Fellow in Hematology, Children's Hospital Medical 
Center, Boston, Massachusetts 
Chief, Clinical Service, Molecular Hematology Branch Chief, 
Section on Clinical Hematology, Molecular Hematology Branch, 
National Heart, Lung, and Blood Institute, National 
Institutes of Health, Bethesda, Maryland 
Deputy Clinical Director, National Heart, Lung, and Blood 
Institute 
Chief, Clinical Hematology Branch, National Heart, Lung, and 
Blood Institute 
SELECTED PUBLICATIONS 
1. Karlsson, S., Humphries, R.K., Gluzman, Y., and Nienhuis, A.W.; 
- Transfer of genes into hematopoietic cells using recombinant DNA 
viruses. Proc. Natl. Acad. Sci. 82:158-162, 1985. 
2. Nienhuis, A.W., Bunn, H.F., Turner, P., Gopal, T.V., Nash, 
W.G., O'Brien, S.J., Sherr, C.J.: Expression of the human c-fms proto- 
oncogene in hematopoietic cells and its deletion in the 5q- syndrome. 
Cell 42:421-428, 1985. 
3. Karlsson, S., Doren, K.V., Schweiger, S.G., Nienhuis, A.W., Gluzman, Y.: 
Stable gene transfer and tissue specific expression of a human globin 
gene using adenoviral vectors. EMBO Journal . 5:2377-2385, 1986. 
Recombinant DNA Research, Volume 16 
[203] 
