ZOOLOGY AND BOTANY. MICROSCOPY, ETC. 
47 
According to Claus. 
The Septal Funnels. 
Claus described the longi- 
tudinal muscles of the Scypho- 
stoma as solid strands, but now 
be recognizes that they arise as 
Goette described, though he 
does not admit the accuracy of 
the term septal funnels, nor the 
homology between septal funnels 
and subgenital cavities. 
The Tentacles and the Radial Symmetry. 
Claus derived the axial part 
of the tentacles from the endo- 
derm of the stomach, and re- 
garded the two primary gastric 
pockets as these axes. The 
majority are interseptal, but the 
four inter - radials are septal ; 
and so on. But Claus has 
now corrected these conclusions, 
adopting those of Goette. 
Claus at first found the 
determination of the radial 
symmetry in the tentacles, but 
he now distinguishes a peri- 
pheral (tentacular) from an 
inner (antimeral) segmentation, 
the latter depending on the 
gastric diverticula. 
According to Goette. 
The longitudinal muscles 
arise in the walls and solid out- 
growths of septal funnels which 
grow from the ectoderm of the 
oral disc. Moreover, these septal 
funnels are really continuous 
and identical with the subgenital 
cavities of the Medusa. 
All the tentacles of the Scy- 
phostoma arise from the gastric 
pouches ; the “ scjital tentacles ” 
arise from the lateral corners of 
the third and fourth gastric 
pouch, and only subsequently 
come to lie over the septa ; the 
contrast which Claus draws be- 
tween the wholly interseptal 
tentacles of Anthozoa and the 
partly interseptal tentacles of 
the Scyphostoma is therefore 
mistaken. 
The symmetry of the Scypho- 
stoma is defined by the four 
gastral pouches, and repeated in 
septa and gastral folds. 
The Morphological Import of the Scyphostoma. 
The Scyphostoma is in its 
structure a hydropolyp with 
gastric folds, and is without the 
essential characteristics of an 
Anthozoon — oesophagus, septa, 
gastric pouches, and only inter- 
septal tentacles. 
The Scyphostoma, until the 
beginning of strobilation, is 
without the essential character- 
istics of a tetrameral or oc- 
tomeral or polymeral Scypho- 
medusa. 
The Scyphostoma is never 
like a simple hydropolyp, or 
even like one provided with 
gastric folds, but it has, in its 
ectodermic oesophagus, in its 
septa, gastric pouches (folds and 
grooves), and interseptal ten- 
tacles, the structure of an 
Anthozoon. 
But this structure lasts only 
for a short time, for in the 
eight-armed stage at latest the 
Scyphostoma is modified into a 
Medusa. 
