Nov., 1912.] The Reduction Division in Fuchsia. 
3 
In his paper on Oenothera lata x O. lamarckiana (5) he finds 
starch grains present in the cytoplasm of the mother cell. These 
grains become more abundant until the reduction takes place 
after which they are not found. He finds what he calls “pro¬ 
chromosomes”, but in a later paper considers these bodies to be 
nucleoli. The presynaptic stages show a continuous spirem 
which is exceedingly delicate and coiled. Then follows the con¬ 
traction, after which follows a stage in which the spirem is much 
shortened and several times thicker than just before the contrac¬ 
tion. He does not believe that the nucleoli break up and move 
out into the cytoplasm waiting to be re-collected into a new 
nucleolus in the new nucleus, as Sehaffner believes to be the case 
in Lilium philadelphicum. There are twelve chromosomes and 
one or two “heterochromosomes”, or as he calls them in a later 
paper—nucleoli. He believes O. lamarckiana to be a pure strain 
and not a hybrid. 
In the paper on Oenothera gigas (8) Gates says “The absence 
or partial absence of a close pairing of chromosomes in diakinesis 
and on the heterotyptic spindle is in strong contrast to the con¬ 
dition in other genera of plants where the chromosomes are 
regularly paired. However * * * similar failure to pair is 
often exhibited. These cases appear to be the exceptions to the 
general law enunciated by Montgomery in 1901 from his observa¬ 
tions on Hemiptera, that homologous chromosomes of maternal 
and paternal origin pair with each other in synapsis. Later 
observations on a variety of forms, in which there are morpholog¬ 
ical chromosome differences, show that ordinarily chromosomes of 
similar size and shape pair with each other and justify the view of 
Montgomery which has been widely adopted. There is some 
variety in the size and shape of the chromosomes but nothing 
constant was observed. ” In the anaphase the longitudinal split 
generally described does not always show. There are some 
irregularities in the reduction such as an unequal number of chro¬ 
mosomes passing to the two poles. 
Davis in his first paper on Oenothera (2) observes the nucleus 
filled with a close reticulum having the chromatin material dis¬ 
tributed around the periphery of the nucleus. Next follows the 
appearance of extremely delicate strands connecting the bodies, 
forming an open net work. These strands readily thicken and 
become more numerous until finally the nucleus is filled with a 
relatively close reticulum. During this time the chromatin 
bodies become smaller and seem to contribute their material to 
formation of the strands so that the only large structures in the 
nuclei are the nucleoli. Synapsis, as he calls the synizetic con¬ 
traction, begins slowly and finally carries the strands away from 
the nuclear wall. During this contraction there is a marked 
change in the structure of the reticulum At first the reticulum 
