Annals of the Transvaal Museum 97 
trochanter is 9 cm. Its thickness, measured from the top of the upper end of 
the trochanter quartus as preserved to the top of the ridge on the anterior 
surface, is 12 cm. 
The section of the bone, which is revealed at the fractured end, is re- 
markable. The fractured surface has 
been polished and its photcfgraph is 
given in fig. 2 of PI. XII. It shows a 
thick wall of substantia compacta 
which is clearly lamellar. Sharply 
defined from this is an inner mass 
of substantia spongiosa. Only a small 
portion of the whole section is not 
occupied by bony matter. Here we 
have, therefore, a Theropod femur of 
practically solid structure. Zittel in 
his “Handbuch” (1) states on p. 718 
that the limb-bones of all Thero- 
poda are hollow and were perhaps 
filled with air. In Eastman’s Text- 
book it is stated in the diagnosis of 
the Sub-Order Theropoda ( 3 , p. 227) 
that the limb-bones are hollow, v. 
Huene, in his great work “Die 
Dinosaurier der europaischen Trias- 
formation,” compares the Theropoda 
with the Sauropoda on p. 346, and 
says: “The limb-bones of the Sauro- 
poda are not tube-like, but nearly 
completely massive, whereas those of 
the Theropoda are hollow.” In the 
second edition of Zittel’s handbook 
in 191 1 , Broili states in the diagnosis 
of the Sub-Order Theropoda that the 
limb-bones are mostly tube-like. 
Probably, therefore, Broili knew of 
Theropod limb-bones which were not 
hollow, and our femur now adds to the evidence. 
Tibia. 
The left tibia (PI. XI, figs. 1 and 2, and text-figs. 6 and 7) is complete. 
It has a very large proximal articulation surface and a straight shaft. The 
bone was found in five pieces, which all fitted neatly together. The total 
length of the bone is 46 cm* The head of the tibia has a length of 19-5 cm., 
measured from the hinder end of the medial condylus to the anterior edge of 
the tuberositas tibiae. Its greatest breadth, transverse to this grea^st length, 
is 13-5 cm. The length of the lateral side of the head is about 15 cm. An- 
teriorly the head of the tibia has a laterally directed process, the tuberositas 
tibiae. The lateral border of the proximal articulation surface is slightly con- 
vex at the tuberositas, more convex around the lateral condylus and sharply 
concave in between. The medial border runs from the tuberositas tibiae in- 
wards and curves gradually backwards; the posterior part of this border is 
not so convex as the anterior portion. The posterior border is convex behind 
each condylus and notched in the middle ; this notch is still filled with matrix. 
Text-fig. 5. Eucnemesaurus fortis. Outline 
of the posterior view of the proximal 
end of the left femur, showing the 
position of the trochanter minor and 
the upper end of the trochanter quartus . 
x 4 . 
