144 
Y. TANAKA. 
It is true that there is no positive reason to deny the possibility of the 
occurrence of intermediate series besides those forwarded by Bateson. But 
we must admit, on the other hand, that a decisive proof for the existence 
of such intermediate systems is also not yet discovered. 15 
Concerning the phenomena of coupling and repulsion there are many 
questions to be answered by futurp investgations. The more important of 
those are : 1) At which stage of the ontogeny of an individual does the 
coupling or repulsion occur?; 2) Is there any selective mating of gametes? 
3.) What is the actual cause of the coupling and repulsion? At present 
I have no data which throws light on these points. 
I trust that I have furnished at least some positive data regarding 
the question whether the phenomena of gametic coupling and spurious 
allelomorphism in animals occur independently of the sex-character. In their 
important paper Bateson and Punnett remark : “Hitherto no case of 
coupling has been found in animals. Among the phenomena of repulsion, 
however, of which many examples exist, certain suspicious cases have been 
observed which may mean that in animals reduplicated systems exist like 
those of plants. Nevertheless at present it seems not impossible that the 
two forms of life are really distinguished from each other in these respects” 
(1911, b). However to-day we have no need to make such a distinction 
regarding animals as distinguished from plants. 
1) Quite recently, Trow (1912) found the 2 : 1 : 1 : 2 coupling between tlie hairness 
and the ray-character in Senecio vulgaris. 
POSTSCRIPT. (March 20, 1913.) Two following data recently described by Hagedoorn 
(The, genetic factors in the development of the Housemouse. Zeits. f. ind. Abst. u. Vererb. Bd. VI, 
pp. 97-136, 1912) and Punnett (Inheritance of coat colour in Rabbits. Journ. Gen. Vol. 2, 
pp. 221-238, 1912) should be added. 
Animal name. 
Characters 
System | 
Author 
Year 
Housemouse. 
A) Colour factor and B) ‘agouti’ factor. 
a) Albino and b) black. 
complete 
repulsion 
Hagedoorn 
1912 
Rabbit. 
A) Deepening- factor for melanic pigment and B) ex- 
tending factor for melanic pigment. 
a) Absence of deepening factor and b) absence of extending 
factor. 
complete 
coupling 
Punnett 
1912 
These are, as I believe, only literature of gametic reduplication between two characters 
independent of the sex-factor in animals. 
