ON Til K TjAMIN AKIACEAK OF HOKKAIDO 
5 
KEY TO THE SPECIES 
I. Blade entire 
A. Mucilage canals present in both stipe and blade 
a. Hapteres not verticillate but mainly in vertical rows 
a. Blade coriaceous 
1. Blade linear-lanceolate, broad, rather round at the base, with the 
insertion of the stipe not in the center of the basal margin . . 
1. L. japonica Aresch. 
2. Blade linear-oblanceolate, usually cuneate at the base 
2. L. ochotensis Miyabe 
/3. Blade fragile 
3. Stipe short (4-6.5 cm), blade narrow (15-20 cm) 
3. L. fragilis Miyabe 
4. Stipe short (6-12 cm), blade broad (ca. 40 cm' 
. . . 4. L. diabolica Miyabe 
5. Stipe long (50-60 cm), blade broad (ca. 52 cm) 
5. L. longipsdalis Okamura 
b. Hapteres verticillate 
6. Blade quite smooth on the surface even while young, with no 
bullations' 0 6. L. religiosa Miyabe 
7. Blade with a row of bullations along both sides of the median 
fascia while young, becoming smooth on the surface and crispate 
on the margins at maturity 7. L. cichorioides Miyabe 
B. Mucilage canals absent from the stipe, present only in the blade 
a. Mucilage canals discontinuous, being scattered here and there; (canals 
rarely present in the stipe too) 
8. Sori on the upper surface 5 6 * * * 10 of the blade along the margins but 
not on the median fascia, and developing from the base upwards 
8. L. angustata Kjellman 
b. Mucilage canals a continuous network 
9. Blade very long, up to twelve fathoms; sori formed from the 
base upwards, on both surfaces . . . 9. L. longissima Miyabe 
5) Bullations are present in L. religiosa while the frond is young. ( rf . YENDO, 
1913, p. 302 ; Okamura, 1936, p. 246 ; Miyabe in Okamura, 1936, p. 284). 
6) Dr. Miyabe (1902, p. 4) stated in his introductory remarks under the heading 
of “External morphology” as follows : “The median fascia is elevated on one surface 
and depressed on the other. The elevated surface is commonly called the upper surface 
whereas the depressed is called the under, because the former is believed to face 
usually the light in the natural state.” However, YENDO (1911, p. 302) describes rightly 
that the elevated surface faces the substratum and the depressed faces the light in 
the natural state, so the latter is to be called the upper surface whereas the former 
the under. 
