14 
K. MI '»'AHE 
Kakijima-kombu (Akkeshi), O-ita-kombu (ditto). 
Holdfast composed of filamentous hapteres divided several 
times. Stipe solid, very long, usually 50-66 cm, sometimes up to 
1 m in length, round in cross section below, complanate and 
broadened upwards. 15-16 mm in diameter near the base. Blade 
lanceolate, broad, cordate at the base when matured but round or 
broad cuneate while immature, slightly undulate on the margins, 
with a median fascia not so distinct as in other species, sometimes 
up to 33-76 cm in breadth, fragile in substance. Sporangial sori 
on both surfaces of the blade, in patches similarly shaped to those 
of Laminaria diabolica. 
The mucilage ducts are present in both stipe and blade ; they 
are smaller in diameter as compared with those of Laminaria 
diabolica. The medulla of the blade is thin, consisting chiefly of 
vertical rows of filamentous cells. 
Habitat. Growing on dead oyster shells heaped in Akkeshi Lake, 
a lagoon at Akkeshi, Kushiro Province. Uuknown from elsewhere. 
Use. Because of its inferior quality, the present alga is not 
fit to be exported. Its annual yield is said to be only 300-400 
koku.’ 10 
Addenda. This species is unique among the Laminaria species 
of Hokkaido in that its occurrence is restricted to a special locality. 
It grows restrictedly in Akkeshi Lake favored by the calm water 
of low salinity, and consequently has quite a long stipe and a 
markedly broadened and thin blade. In other important characters, 
however, it bears a close resemblance to Laminaria diabolica, which 
grows in Akkeshi Bay at places only a few kilometers distant from 
the habitat of L. longipedalis. In view of this fact, there may be 
a question as to whether it is more reasonable or not to recognize 
the latter species as belonging to a variety of the former. 1 ' 0 The 
author describes it as a distinct species here following the view 
of Dr. K. Okamuka, who named it Laminaria longipedalis. 
15) One koku is equivalent to 330.6 lbs. 
16) MlYABE and Nagai (1933, p. 88) reduced the preceding described species 
to a variety of the present, as var. diabolica MlYABE, according to the rule of priority, 
and Okamura (1936, p 250) agreed with their view. However, Miyabe [in Okamura, 
1936, pp. 285-286 treated it again as a distinct species. 
