72 
Psyche 
[March 
petiolation but of this we have no information. Bolton figured and 
described the costal margin at the level of the middle of the petiole 
as having a distinct hump-like elevation but I have not seen this in 
any of the specimens which I have examined. There is some uncer- 
tainty about the shape of the apex of the wing. Curiously enough, 
this is missing in all specimens of the species which have been indi- 
vidually described; none of those which were described and figured 
by Handlirsch and by Bolton show the apex; Bolton includes the 
apex in his diagrammatic figure of the complete wing but he does 
not indicate the specimen on which the drawing was based. How- 
ever, the shape of the wing apex as shown in his “complete wing” 
is more broadly rounded than that in any specimens which I have 
seen or which have been described by Handlirsch. The wing rep- 
resented by Laurentiaux (1953) in an original drawing based upon 
specimens in the Birmingham Museum has a much more bluntly 
rounded apex; however, his entire figure seems to be somewhat 
diagrammatic and is probably not intended to represent the precise 
form of the wing. The absence of the apical part of the wing in so 
many specimens suggests that the wings of Brodia were unusually 
fragile and delicate. 
(2) The serration of the wing margins. The serrate margin is 
one of the more obvious features of Brodia and it has been figured or 
mentioned in virtually all accounts of this species. The description 
and especially the figure given by Scudder in 1883 are accurate and 
detailed. The structure of the costa is very clearly preserved in the 
specimens in the M.C.Z. (Scudder collection); they show that the 
costa has the serrations along both its anterior and posterior edges, 
those of the anterior edge being somewhat sharper. Beyond the point 
where the subcosta joins the costa the serrations along the posterior 
margin of the costa become more pronounced, as correctly drawn by 
Scudder (see top fig., Plate 14). The subcosta, which is not actually 
fused or united with the costa, remains visible as an independent 
vein. In one of the specimens in the Scudder collection the costal 
margin is very clearly preserved and shows also in the distal part 
serrations or short spines on the subcosta in that region, making a 
total three rows of these short projections. 4 It is to be noted in this 
connection that the radius also bears a few setal bases (see Plate 14). 
The serrations become very small along the posterior margin distally 
4 The costal structure of Brodia resembles to a striking degree that of the 
anisopterous Odonata, in which the costa also has serrations on both the 
anterior and posterior borders. 
