IO 
Psyche 
[March 
The two workers are very similar in most characters, but differ 
markedly in size. Both are roughly what would be called “medium- 
sized” ants. Some exact measurements could be taken, to the nearest 
0.0 1 mm, and these are given in millimeters as follows: 
Specimen i. Head width not measured ; head length along mid- 
line 0.98 ; maximum length of right eye 0.32. Segments of left an- 
tenna (numbered I through XIL) from insertion outward: I (scape) 
0.60, II 0.16, III 0.34, IV 0.21, V 0.21, VI 0.21, VII 0.21, 
VIII 0.21, IX 0.21, X 0.21, XI 0.21, XII 0.26. (Holotype). 
Specimen 2. Head width taken just anterior to the compound 
eyes, 1.00; head length not measured. Segments of right antenna 
(numbered I through XII) from insertion outward: I (scape) 0.68, 
II 0.18, III 0.40, IV 0.27, V 0.27, VI 0.26, VII 0.26, VIII 0.26, 
IX 0.24, X 0.23, XI 0.23, XII 0.31. (Paratype). 
The details of morphology, including pilosity, are evident in the 
figures. Pubescence consists of scattered, short (about 0.01 mm long), 
appressed hairs. Both specimens are uniform light brown in color, 
but this means little in view of the passage of 100 million years. 
Holotype: worker, labelled Fi, shown in photograph, Plate 1 ; 
paratype: worker, labelled F2, shown in photograph, Plate 3. Both 
specimens were collected in a single piece of amber (now in sep- 
rate pieces) in clay, Cliffwood Beach, on Raritan Bay, New Jersey, 
by Mr. and Mrs. Edmund Frey. The specimens are in the collection 
of Mr. and Mrs. Frey, Mountainside, New Jersey. 
Phylogenetic position of Sphecomyrma 
Sphecomyrma freyi presents a mosaic of wasp-like and ant-like 
character states. Moreover, most of the ant-like features are, as 
well as we can judge the matter, primitive with respect to the other 
known Formicidae. Our assignments of various character states 
are given in Table 1. An examination of this arrangement will show 
that Sphecomyrma is truly intermediate between the primitive ants 
Figure 3. Alitrunk of Methocha malayana female in side view. (Re- 
drawn after Pagden, 1949). 
