1967] 
Barr — New Pseudanophthalmus 
169 
width, which occurs at apical fifth; base a little more than 9/10 as 
wide as apex; margins strongly rounded in apical half (in apical 
third in P. fuscus) , then oblique, subparallel in basal sixth; hind 
angles prominent, right, sharp, and reflexed, the paramedian foveae 
deep; base feebly emarginate between hind angles. Elytra elongate- 
elliptic, length 1 3/5 times combined width; prehumeral borders 
slightly oblique to mid-line, humeri somewhat angular but rounded, 
humeral margins setose but not serrulate; disc with 5 or 6 ir- 
regular rows of pubescence on each interval ; longitudinal striae 
broadly and very shallowly impressed (obsolescent in P. fuscus ) ; 
distance between 1st and 4th umbilicate punctures 4/5 the distance 
between 4th and 5th punctures (3/5 in P. fuscus) ; anterior discal 
puncture at level of 4th umbilicate puncture, both anterior and pos- 
terior discals in 3rd stria; recurrent portion of apical groove short, 
rounded to subparallel, bending sharply inward to apex of 3rd 
stria at its anterior terminus; anterior apical puncture posteromedial 
to juncture of recurrent groove and 3rd stria. Appendages : antenna 
5/1 1 the total body length, outer segments rather short and thick, 
9th segment 3/5 as wide as long (1/3 in P. fuscus ), densely pubes- 
cent; mentum tooth broad with apex emarginate; legs long and 
slender, but proportionately less slender than in P. fuscus. Male 
with secondary sex characters normal. Aedeagus 0.74 mm long 
(holotype) (0.88-0.90 mm in P. fuscus) ; basal bulb moderately 
expanded, with constriction before median lobe; apex gradually at- 
tenuate and very finely knobbed, not produced and conspicuously 
knobbed as in P. fuscus ; parameres moderately elongate, with 3 apical 
setae, the middle one shortest; internal sac virtually unarmed, cop- 
ulatory sclerites db tabulate as in P. fuscus but shorter. 
Holotype cf (United States National Museum) and 2 99 para- 
types, east base of Kennison Mountain near the Cranberry Glades, 
elevation 1000 meters, Pocahontas Co., West Virginia, 28 April 1967 
(T. C: Barr, leg.). Measurements of holotype: total length 4.12 
mm, head 0.72 mm long X 0.74 mm wide, pronotum 0.72 mm long 
X 0.96 mm wide, elytra 2.22 mm long X 1.40 mm (combined) 
wide, antenna 2.32 mm long, aedeagus 0.74 mm long. 
The discovery of P. sylvaticus strongly favors the hypothesis ( Barr, 
1965) that the ancestors of Nearctic cave trechines had already 
undergone much eye and pigment reduction and become edaphobites 
(obligate inhabitants of soil and humus) before colonizing caves. 
Under a periglacial climate, trechines would have been much more 
widely distributed than at present. Following retreat of the glaciers, 
