1967] 
MacLeod & Adams — Berothidae 
243 
sternite, in some cases very large (cf Nosy bus) , is present in nearly 
all berothids and is usually bilobed it seems unlikely that such an 
alternative interpretation is correct. 
The only other Berothidae described from southern South America 
are Naizema patagonica Navas and Espetera mendozina (Esben- 
Petersen). Neither of these has been adequately known and because 
of the presence of recurrent humeral veins in the wings of N. pata- 
gonica, , doubt has been expressed that this species is even a berothid 
(Carpenter, 1940; Tjeder, 1959). While it has not been possible 
to examine Navas’ type of N. patagonica , male and female cotypes 
of E. mendozina have been loaned to us for study. Examination of 
these specimens and of Navas’ descriptions of N. patagonica (1919, 
1929) has convinced us that Esben-Petersen’s species is certainly con- 
generic with N. patagonica and, indeed, is very likely the same species. 
Although it shows no close similarity to Cyrenoberotha , a redescrip- 
tion and catalogue of Naizema will be presented here as the rather 
unusual structures of this group provide insight into some of the 
structural specializations of the Berothinae. 
Naizema Navas 
Figures 20-25 
Naizema Navas, 1919, Rev. R. Acad. Cienc. Madrid 17: 298-299. 
Type species (by original designation) : Naizema patagonica Navas. 
Navas, 1929, Mem. Acad. Cienc. Zaragoza 2: 19-20. Handschin, 1935, 
Rev. Suisse Zool. 42: 700. Carpenter, 1940, Proc. Amer. Acad. Arts 
Explanation of Plate 32 
Cyrenoberotha penai n. sp.: Figs. 7-9 of $ holotype. Lomamyia squamosa 
Carpenter, $ : Figs. 10-13, drawn from a specimen from 3 mi. NW of La 
Palmilla on Highway Sur No. 19, B. Calif., Mexico; 18-1-59; H. B. Leech 
Col.; in the collection of the California Academy of Sciences. This specimen 
is the same as that studied and figured by Acker (1960), where it is identified 
as “ Lomamyia sp.” 
Fig. 7. Lateral aspect of terminalia. Fig. 8. Detail of gonarcus, gonocoxites, 
mediuncus and hypandrium internum, drawn after the removal of these 
structures from the abdomen, lateral aspect. Fig. 9. Same as Fig. 8, pos- 
terior aspect. Fig. 10. Divided dorsal and ventral views of fused gonarcus 
and gonocoxites and of mediuncus (terminal tuft of setae of mediuncus 
omitted), inset showing detail of microtrichia on surface of gonarcus. Fig. 11. 
Same, lateral view. Fig. 12. Ventral view of mediuncus, inset showing detail 
of tips of two longest (median) pairs of setae. Fig. 13. Lateral view of 
terminal abdominal segments of abdomen. 
Abbreviations: ect — ectoproct; gcx — gonocoxite; gcx. ap. — basal apo- 
deme of gonocoxite; gs — gonarcus; hy. i. — hypandrium internum; mu — 
mediuncus; 9S — 9th sternite; 8T, 9T — 8th, 9th tergites; 9T + ect — fused 
9th tergite and ectoproct. 
