1967] 
MacLeod & Adams — Berothidae 
251 
of only a small number of South African species, Tjeder’s divisions 
seem quite sound, although, as noted below, we believe that several 
of the genera which he referred to the “Sphaeroberothinae” (on the 
basis of their rounded wing tips) should be reassigned to the Ber- 
othinae. In addition it is now necessary to propose a new subfamily 
for the inclusion of Cyrenoberotha. 
In the following section we have provided new descriptions for 
the four subfamilies and have listed all of the described genera. We 
have had representatives of all of the subfamilies and principal genera 
available for this study, the specific genera we have examined being 
marked with an asterisk in the list. Other than the synonymy of 
Espetera and Naizema noted above, all described genera are listed 
without indication of synonymy since so few of the type species have 
been examined by modern workers. Our views on certain obvious 
synonymies are set forth in the discussions following the subfamilies. 
The four subfamilies of the Berothidae may be distinguished by 
the following key: 
1. Forelegs cursorial; free stem of MA in hind wing vertical, 
straight 2 
Forelegs raptorial; free stem of MA in hind wing oblique, 
sinuate Rhachiberothinae 
2. Ninth abdominal tergite fused with ectoprocts; face very short, 
not much extended ventrad of lower margin of eyes 3 
Ninth abdominal tergite free from ectoprocts; face conspicuously 
extended ventrad of lower margin of eyes . ... Cyrenoberothinae 
3. Bases of R and M fused in forewing, their common stem showing 
no trace of a composite structure, with MP diverging abruptly 
from R + MA at an angle exceeding 30° ; 8th sternite of 
female an elongate, triangular sclerite extending posteriorly 
beneath the fused 9th tergite and ectoprocts and the bases 
of the similarly produced 9th gonocoxites; hypocaudae lack- 
ing Nosybinae 
Bases of R and M in forewing usually separate, where closely 
adajacent or fused these bases usually preserving some trace 
of their separate origins and with MP diverging from R + 
MA at a more acute angle which is usually less than 20° ; 8th 
sternite of female not greatly elongate ; hypocaudae present 
(figs. 18, 23) or absent Berothinae 
Cyrenoberothinae n. subfam. 
Description. Berothids with head and mouthparts elongate; ver- 
