258 
Psyche 
[September 
Navas, 1929 (Australia) ; Dasypteryx Stein, 1863 (Greece) ; *Es- 
petera Navas, 1929 [= Naizema Navas, 1919, see above] (Ar- 
gentina) ; *Frawalkeria Navas, 1929 (Borneo, Formosa); 
Isocelipteron Costa, 1863 (southern Europe) ; Lekrugeria Navas, 
1929 (India) ; '* Lomamyia Banks, 1904 (North America, we have 
also seen a specimen from Zamorano, Honduras) ; Naizema Navas, 
1919 (Argentina) ; *Nodalla Navas, 1927 (Egypt) ; Podallea Navas, 
1936 (Ethiopia) ; Protobiella Tillyard, 1923 (New Zealand) ; 
Sisyrura Navas, 1905 (China) ; *Spermophorella Tillyard, 1916 
(Australia) ; *Sphaeroberotha Navas, 1930 (Tunisia) ; *Stenobiella 
Tillyard, 1916 (Australia) ; *Trichoberotha Handschin, 1935 (Aus- 
tralia) ; Trichoma Tillyard, 1916 (Australia). 
Kruger (1923) has also described a species, Proberotha prisca , 
from the Baltic amber; unfortunately this specimen has not been lo- 
cated for study so that its subfamily placement cannot now be 
ascertained. An undescribed amber berothid in the collection of the 
Museum of Comparative Zoology which, judging from Kruger’s 
rather incomplete description of P. prisca , is probably not congeneric, 
has long hypocaudae and is obviously a berothine. 
The genera Dasypteryx , Frawalkeria, Isoscelipteron, Podallea , and 
Sisyrura are obviously very close to Berotha and several of these 
have long been regarded as synonyms of this genus. Lekrugeria and 
Acroberotha, although presumed to differ from Berotha in the num- 
ber of branches of the radial sector of the forewing and shape of the 
apex of the wings, are also at best only poorly distinct and may 
prove to be synonymous with Berotha when the structure of these 
species becomes known in detail. An appreciation of the variability 
of such characters as the branching of the radial sector and the shape 
of the wing apex can be gained from a review of the species of the 
well-studied genus Lomamyia. These show considerable individual 
variation in the number of branches of Rs, reaching as high as 7 (not 
counting MA) in some specimens of L. fulva Carpenter, and present 
wing shapes ranging all the way from slightly pointed (L. longicollis 
(Walker) ) to species with strongly falcate apices ( L . hamata 
(Walker), L. squamosa Carpenter). It seems unlikely that stable 
generic limits will be found utilizing such characters as these. 
Notes on the evolution of the berothidae 
The Subfamilies of the Berothidae. An estimate of the overall 
degree of evolutionary change from the presumed structure of an- 
cestral berothids of eight key genera of the Berothidae is presented 
