3 1 8 
Psyche 
[December 
(1929) describes the condition in some other species. Of previously 
known Oriental Siagona, only crassidens (of southern Indo-China) 
has the male mandible armed nearly like that of the present new 
species, but the latter, compared with Bates’ (1889) description of 
crassidens , is larger, with head ridges more abbreviated posteriorly, 
with sparser elytral punctation, and with the mandibular tubercle 
not bilobed. 
Andrewes’ key (referred to above) is based chiefly on asexual 
characters and does little to help place the new species. The latter 
does not fit into the key: it is winged and large (couplets 1 and 2), 
but the lateral ridges of the head do not nearly reach the neck con- 
struction. In wings, size, and head ridges the new species is like sub- 
laevis Ghaudoir, which is known only from southern Indo-China, 
Siam, and the Malay Peninsula, and which is therefore omitted from 
the “British India” key. There are two specimens, both females, of 
sublaevis in the Museum of Comparative Zoology, and the female of 
the new species matches them in most characters including the pres- 
ence of an unusually distinct, short, post-humeral plica on each ely- 
tron. The only significant difference I note is sparser punctation of 
the disk of the elytra in the new species. But Chaudoir (1876) 
described sublaevis f from Bangkok, from a good series of both sexes, 
and the male has no mandibular tubercle. The Javan insulana An- 
drewes (1936), of which I have one male, i:s a smaller species, with- 
out distinct post-humeral elytral plicas, but with nearly similar head 
ridges. It lacks mandibular tubercles. 
Taking one consideration with another, I think that the new 
Philippine species most likely represents the stock of sublaevis, and 
that the mandibular tubercle of the male has been developed inde- 
pendently, not inherited from an Asiatic ancestor, and I suspect that 
insulana may prove to be a modification of the same stock rather 
than a relative of fabricii Andrewes as Andrewes (1936) originally 
supposed. It will require a revision of a number of Oriental Siagona 
to decide these possibilities. 
References 
Andrewes, 
1929. Fauna of British India etc., Coleoptera, Carabidae I. 
1933. Annals and Magazine of Natural History (10)11: 99. 
1936. Annals and Magazine of Natural History (10) 17: 307-308. 
Banninger 
1918. Deutsche Ent. Zeits. for 1918: 97-109. 
