400 
Psyche 
[Vol. 92 
ouridae, Taeniopteryidae and Perlodidae. Ephemeropterans included 
representatives from the genera Ephemerella, Epeorus, Cloeon, 
Stenonema and a member from the family Baetidae. Trichopterans 
included representatives of the genera Glossosoma, Hydropsyche 
and Cheumatopsyche. 
Results of monthly foregut analysis are presented in Figure 4. 
Plecopterans and ephemeropterans made up the principle compo- 
nents of the diet throughout the year. Dipterans became important 
in the winter and summer, while trichopterans were most important 
in the summer. Naiads fed most during the winter months with an 
average of 5.0 food items/ stonefly. They fed least during the spring 
and fall with an average of 1.1 food items/ stonefly each season. 
Siegfried and Knight (1976a, 1976b) and Sheldon (1969) reported 
dipterans, trichopterans and ephemeropterans as the most impor- 
tant food items for C. calif ornica. Sheldon (1969) found that ephem- 
eropterans were a regular component of C. californica’’ s diet and 
dipterans were important in the winter and summer. Siegfried and 
Knight (1976a) stated that dipterans were the most important and 
consistent food item for C. californica. Siegfried and Knight (1976b) 
found that C. californica naiads showed no preference for a particu- 
lar prey and would eat anything available. They also reported a 
significant correlation between gut diversity and benthic diversity. 
Sheldon (1969) stated that food consumption was low in the 
summer. In a laboratory study, Heiman and Knight (1975) found 
that C. californica had its greatest food consumption in the summer, 
with minimal feeding in the winter, at normal environmental 
temperatures. Johnson (1981) reported that dipterans and trichop- 
terans were the primary diet for A. abnormis. The diet was supple- 
mented with ephemeropterans. 
Phoresy. Larvae of a chironomid midge were observed living 
phoretically with Acroneuria carolinensis naiads. The midges 
occurred on the prothorax or just beneath the mesothoracic wing 
pad. The phoretic attachment occurred on nine of 241 (3.7%) naiads 
examined. Dosdall and Mason (1981) reported the chironomid 
Nanocladius ( Plecopteracoluthus ) branchicolus living phoretically 
on A. lycorias. They reported varied percentages of the number of 
stoneflies bearing the chironomids. The percentages of phoretic 
cases ranged from 10-50 percent. 
Rearing and emergence. The attempt to collect emerging adults 
on 31 May 1984 was unsuccessful. Exuviae were collected up to 4.6 
