136 
Psyche 
[Vol. 93 
much of interantennal crest), except for brown on antennal sockets, 
brown also on a large to very large, quadrangular to (more often) 
dorsally narrowed median facial blotch (which is occasionally 
reduced to a small pale brown tinge and which sometimes, when 
conspicuous, surrounds a yellow area along clypeo-frontal suture), 
and also brown on a large to small or even obsolete area in and 
(frequently) above and below anterior tentorial pit, which may be 
confluent dorsally with the median brown facial area; sometimes 
also with yellow in malar space and broadly bordering hind orbit to 
as much as upper 0.2 of eye; and yellow also on most of clypeus 
except for its pale brown apical margin (clypeus rarely in large part 
brown with yellow only laterad); on most of basal 0.7 of mandible; 
on maxillary palpomeres 1-3; sometimes on an anterio-ventral fore 
coxal blotch; occasionally on a small dorso-lateral mid coxal blotch; 
on a broad anterio-dorsal stripe on fore and sometimes mid tro- 
chanters (yellow on mid trochanter often dull and weakly de- 
veloped); sometimes also anterio-dorsally on fore and mid trochan- 
telli; on a broad anterio-dorsal front femoral stripe; and rarely also 
on part of mid femur anterio-dorsally. 
Length of fore wing: 14.6-18.5 mm. Malar space: 0.63-0.71 as 
long as basal width of mandible. Hind tarsus: segments 1-4 beneath 
with setae longer and denser than in female, pale gray, obliquely 
outstanding, closely packed, 0.4 as long as depth of tarsomeres. 
Clasper: in lateral view with dorsal margin on apical 0.46 broadly 
concave; dorso-apical angle semi-acute (not spiniform) and slightly 
upcurved; apical margin reclivously oblique; apico-ventral angle 
blunt. Other characters as described for female. 
Type Material. Holotype $\ COSTA RICA, Guanacaste Pro- 
vince, Santa Rosa National Park, D. H. Janzen, 1984 (Washing- 
ton). Paratypes: 135 and 9$\ same data as Holotype: 2$ (Wash- 
ington), 1? and 1(5 (Cambridge), 19 (College Station), 1$ and 1<5 
(Gainesville), 1$ and \$ (Lawrence), 1$ and 1(5 (London), 1? and 
1(5 (Los Angeles), 19 and 1(5 (New York), 1? and \$ (Ottawa); 1? 
and 15 (Philadelphia); 1$ and 1(5 (Townes); 1? (Porter). 
Variation. Thyreodon santarosae shows unusually marked 
intrapopulation variability in wing and leg color. This variation 
correlates appreciably but imperfectly with sex. Of the 139 exam- 
ined, 10 have the wings predominantly yellow and in 9 of these 
