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[Vol. 93 
the rest of the wing. Banks (1943) stated that the apical lobes of 
segment X are longer in O. unicolor than in the other species; in 
females of the unicolor complex that we have examined these lobes 
are elongate but variable, and discontinuously longer in none. In the 
holotype there is a distinct tooth or ledge at the base of each lobe of 
segment X on the lateral margin (e.g. Fig. 6); expression of this 
character also shows continuous variability in our material and the 
ledge is lacking in most specimens (e.g. Fig. 3a). The median and 
lateral vulval lobes of the holotype taper to rounded points, which 
blend continuously with a range of conditions in our material. 
The male has not been clearly identified in the literature. The 
illustration of male genitalia labeled as unicolor by Ross (1938, fig. 
48) was given only the status of the “supposed male oi ‘unicolor"' by 
Banks (1943); and although no locality data were given for the 
specimen illustrated by Ross, Banks (1943: 364) stated that it came 
from Inyo Co., California. Banks himself (1943) referred to speci- 
mens from Banff and Alaska that “may be males of this species,” 
offering as a diagnostic character that the third and fourth spines of 
the paramere are not widely separated. Our material shows such a 
very wide range of variation in arrangement of the spines of the 
parameres that this character cannot be regarded as distinctive. 
O. tristis (Banks). We have examined the three specimens 
(South Park, Colorado) in the type series from the Museum of 
Comparative Zoology. From these specimens Ross (1938) desig- 
nated a lectotype male (17 Aug. 1899) and lectoallotype female (20 
Aug. 1899); the remaining female (20 Aug. 1899) is identical to the 
lectoallotype. Ross (1938) placed O. tristis in synonymy with O. 
unicolor, but Banks (1943) maintained that the two were distinct 
species. Although the females in the type series were characterized 
by a pronounced basolateral tooth or ledge on segment X (Banks 
1943), there seems little difference between these specimens and 
what remains of this character in the holotype of O. unicolor (see 
above). The apices of the posterior lobes of segment X are closely 
appressed in both females of the type series, and all three parts of 
the vulval lobe are truncate. This latter character contrasts with 
somewhat more rounded lobes in the holotype of O. unicolor, but 
we have many specimens showing intermediate conditions. The 
male in the type series of O. tristis was distinguished by narrower 
external branches of segment X (superior appendages of Banks), but 
