1986] Wiggins & Richardson — Onocosmoecus 201 
parameres. The external branches of segment X (superior appen- 
dages of Banks) are slightly narrowed at the base, but this feature is 
variable and appears not to be of diagnostic value. The two corne- 
ous points on the fore wings are darkly pigmented and each is 
surrounded by a fairly well defined dark area. Within the material of 
the unicolor complex that we have examined none of these charac- 
ters is distinctive, and we find no reliable basis for identifying this 
species. 
O. flavus (Martynov). Recognition of this Palaearctic species 
was somewhat irregular in that the description of the female 
appeared as Dicosmoecus sp. (sp.n.?) (Martynov 1913: 477), with 
the name proposed later (Martynov 1914: 253). To the original 
description, Martynov (1913: 478) added the comment: “This spe- 
cies resembles D. unicolor Banks from Washington Territory. But 
having seen no specimens of the last named species, and the struc- 
ture of its genital appendages being entirely unknown, I cannot 
identify my specimens with D. unicolor Judging by the illustra- 
tions, the female appears to have been described again as Dicosmo- 
ecus sp. (Martynov 1925, figs. 1, 2). The male was described and 
illustrated by Ulmer (1927) along with the female; Ulmer mentioned 
the surprising similarity between flavus and the North American 
coloradensis [= unicolor ] which he had described earlier. We have 
examined from the Zoological Institute, Academy of Sciences, 
Leningrad, one of the two female syntypes (Pushino, Kamchatka 
R., 19 July 1908) and a male evidently identified by Martynov; and 
in the ROM are additional specimens from two localities in Kam- 
chatka (Dalneje Lake, 1(5, 19, and Ponomarskaya R., 1(5), and 
from the Kurile Islands (Chishima, 1(5) (Fig. lc). The syntype 
female (Fig. 7) fits readily into the range exhibited by our Nearctic 
material, and we found no unique genitalic characters; the dorsal 
lobes of segment X lack a basolateral tooth and the median vulval 
lobe is narrow and well separated from the lateral lobes. In genitalic 
characters this flavus syntype is very close to females from Washing- 
ton (Olympic National Park, 29 June-1 July 1969, ROM #690148) 
and from the Yukon (Dempster Hwy., km. 72, 1 Aug 1979, ROM 
#791 191b); other Washington females (Minotaur Cr., Chelan Co., 
Sept. -Oct. 1976, S. D. Smith coll.) have several similar characters. 
By contrast, in the Dalneje Lake female, segment X has a basolat- 
eral tooth and the median vulval lobe is broad with its sides largely 
