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[Vol. 93 
eight km. distance. The eastern extent of the viator range is delim- 
ited by the contact of the St. Louis/ Salem and Warsaw limestones 
and, perhaps more directly, by the absence of H. subterraneus 
further east (Barr, 1979b). As is the case with nominate tellkampfi, 
populations of viator are known from caves on both the north and 
south sides of the Green River. The northern limits of the viator 
range are set in large part by a sandstone ridge and extensive fault 
zone across Hart County. This geological feature also appears to be 
a complete barrier to gene flow between the northern subspecies N . 
t. henroti and either nominate tellkampfi or viator to the south (Fig. 
1) (Barr, 1979b). Despite the absence of any known hybrid popula- 
tions, tellkampfi and henroti are the most similar subspecies 
morphologically, and henroti also shows a large degree of morpho- 
logical affinity with viator as well (Barr, 1979b). 
Previous studies using gel electrophoresis (Giuseffi et al., 1978; 
Turanchik and Kane, 1979) have shown that genetic variability in 
local populations of N. t. tellkampfi approach those observed in 
similar surface dwelling invertebrates. These results, coupled with 
similar subsequent findings in other species (e.g., Dickson et al., 
1979), suggest that cave adaptation does not necessarily result in a 
reduction in genetic variation. Further, genetic similarity values (I) 
(Nei, 1972) among eight local populations of nominate tellkampfi 
fall in the range (i.e., 0.90-1.00 (Turanchik and Kane, 1979)) com- 
monly reported for populations of continuously distributed surface 
dwelling species. These results substantiate the contention that con- 
tinuous limestone expanses can act as underground dispersal high- 
ways for cave limited species (Barr, 1968). 
The purpose of the present study was to examine electrophoreti- 
cally several local populations of each of the other three subspecies 
of N. tellkampfi. We were interested in determining how infrasub- 
specific variation in these subspecies compared with that of nomi- 
nate tellkampfi. Further, we wished to use these electrophoretic data 
to quantitatively assess relationships among subspecies and also to 
gain some insight to how the present distributional pattern of the 
species has been produced. In these regards, Barr’s (1979b) morpho- 
logical and biogeographic work provides a model against which the 
electrophoretic data can be examined. 
