1986] 
Brady — Ne arctic Gladicosa 
303 
than gulosa (compare Table 1 with Table 2) and is usually darker in 
color with a more distinct pattern (compare Fig. 4 with Fig. 3). In 
most specimens of pulchra the venter of the abdomen is dark brown 
to black behind the epigastric furrow, while that of gulosa is 
yellowish to light brown. Differences between female and male 
genitalia of these two species are noted under gulosa and in the keys. 
Natural History. Little is known of the habitat or behavior of 
pulchra. I’ve collected this species in Florida from the trunks of 
deciduous trees where their color blends well with the bark sub- 
strate. G. B. Edwards (personal communication) has collected spec- 
imens from similar microhabitats in Florida. Pat Miller (personal 
communication) reported collecting both male and female pulchra 
from the trunks of pine trees at night in Perry, Florida, on 
December 5, 1982. Montgomery (1904) reported finding pulchra 
near Austin, Texas, in drier habitats than gulosa and less 
abundantly. He noted that the females live under stones where they 
make a shallow horizontal burrow lined with silk. Whether this 
behavior is consistent throughout the life cycle or represents a 
temporary adjustment to molting or egg laying is a question to be 
answered. Gladicosa pulchra is not the abundant inhabitant of 
deciduous leaf litter, as are gulosa and huberti. Of the species 
investigated pulchra is the most variable in coloration of the body 
and structure of the epigynum. It is possible that more than 
one species is represented in this complex. 
Roble (1986) reported rearing Mantispa viridis from a Gladicosa 
pulchra egg sac. It is the first record of a lycosid spider serving as a 
host of M. viridis. When the spider died, its egg sac was opened and 
a mantispid cocoon and 95 surviving spiderlings were found. This 
corroborates an earlier observation of high spiderling survival 
within a mantispid-infested egg sac of Lycosa rabida. 
Distribution. From Long Island, New York, along the East 
Coast to Texas in the southwest. Limited in its northern range 
inland to the southern parts of Kansas and Missouri and northern 
Kentucky. More abundant in the southeastern United States (Map 
2 ). 
Records. UNITED STATES. New York. Suffolk Co.: Coram, 
Long Island, 19 Oct. 1934, $ (E. L. Bell). New Jersey. Cape May 
Co.: Cape May, 29 Sept. 1945, $ (C. & M. Goodnight). Virginia. 
Alexandria (Indep. City); Falls Church (Indep. City); Fairfax. Ken- 
tucky. Woodford Co.: Kentucky River, 16 Sept. 1920, $. Tennessee. 
