1986] Luykx, Michel, & Luykx—Incisitermes 353 
colonies were distributed essentially at random (Luykx et al., 1987), 
and will not be further considered here. 
Results and Conclusions 
The distribution of the castes in nine pieces of wood is represented 
in Fig. 1. In eight of the nine pieces a single colony was found. In 
one piece two colonies were found: PL487, and a small incipient 
colony consisting only of the royal pair, one soldier, one larva, and 
five workers. This small colony, PL488, contained entirely within a 
single short segment, does not, of course, give any information on 
caste distribution, and will not be considered further. 
The major portion of most of the colonies (with the exception of 
PL476 and PL486) was found toward the bottom, where the wood 
was less deteriorated and less fragile. (The topmost portions of the 
dead trunks are often thoroughly tunneled and in a highly deterio- 
rated condition, and rarely contain any termites.) In most colonies, 
the king and queen were found together in the lower part of the 
colony (Fig. 2). It seems likely that the royal pair might initiate the 
colony at any level in suitable dead wood, but then move down into 
sounder wood as the colony grows. 
Larvae (the first two or three instars) were found preferentially in 
the same segments as the royal pairs. As illustrated in Fig. 2, among 
the segments with reproductives, 7 out of 9 had more larvae than 
expected for those segments. Twelve of the 14 segments with more 
larvae than expected also had a reproductive or was adjacent to one 
that did. That this is a real association, and not just a common 
tendency for both larvae and reproductives to be located in the 
lower parts of the colony, is suggested by colony PL476, the one 
colony in which the reproductives were found in a segment in the 
upper part of the colony: in this colony the larvae also were concen- 
trated in this same segment (Fig. 2). 
The members of the different castes representing successive stages 
of development — workers, early-stage nymphs, late-stage nymphs 
and alates — showed successively greater degrees of aggregation. For 
example, when the cumulative proportions of workers, early-stage 
nymphs, and late-stage nymphs in colony PL482 are plotted separ- 
ately as a function of the segment number in which they were found, 
it is apparent that the workers were distributed over a wider number 
of segments than the early-stage nymphs, and the early-stage 
