88 Psyche [June 
cidse verse/’ of the “Myrmicidse” but mentioned its affinities 
with the “Dorylides.” 
Forel in 1893 did not mention the genus but might have 
meant to include it in the subfamily Myrmicinse, when he said 
that the fourth tribe (“Myrmicii”) included “les autres genres de 
la sous-famille des Myrmicinse.” And later (1901) he excludes 
it from the Dorylinse when he says, “Done, je maintiens la sous- 
famille Dorylinoe limitee aux genres Dorylus, Aenidus, Eciton et 
Cheliomyrmex.” In von Dalla Torre’s “Catalogus” (1893) it 
was still in the Myrmicinse but near Trigonogaster and Pheidolo- 
geton. In 1895 Emery was still of the opinion that it belonged in 
the subfamily “Myrmicini” in the second tribe (“Myrmicii”) 
near Huberia and Phacota. But nine years later (1904), after 
describing the female of L. revelierei Emery, he returned it to its 
original subfamily (Dorylinse). In the following year, however, 
in Ashmead’s skeleton it stood between “?Liomyrmex” and Epi- 
pheidole in the tribe Stenammini, subfamily “Myrmicinse,” 
family “Myrmicidse”. In 1907 Santschi described males of 
three species, which he referred to the Genus Leptanilla and 
claimed that their doryline affinities justified Emery’s original 
allocation of the genus. It should be noted, however, that 
males of Leptanilla have never been taken with females or work- 
ers; hence it is not certain that those described by Santschi 
belong to this genus. In the “Genera Insectorum” (1910) Emery 
established for the genus a separate tribe (Leptanillini) in the 
subfamily Dorylinse, where it seemed destined to abide in isola- 
tion; Wheeler (1910: “Tribe Leptanillii”), Forel (1917), and 
Forel (1921: tribe not given), and Wheeler (1922) have not dis- 
turbed it. But recently Wheeler (1923) has suggested that even 
further isolation may be necessary: 
“Most myrmecologists recognize only five subfamilies of 
ants and regard the Cerapachyinse as belonging to the Ponerinse, 
the Pseudomyrminse to the Myrmicinse. It is probable, how- 
ever, that future myrmecologists will increase the number of 
subfamilies. I believe that the tribe Leptanillini, which Emery 
includes among the Dorylinse, will have to be separated out as 
a distinct subfamily (Leptanillinse). Dr. George C. Wheeler 
