1940] 
The Germs Echestypus 
85 
NOTES ON HIPPOBOSCIDiE. 
14. THE GENUS ECHESTYPUS SPEISER 
By J. Bequaert 
Department of Comparative Pathology and Tropical 
Medicine, Harvard Medical School, Boston, Mass. 
Echestypus Speiser 
Echestypus Speiser, 1907, Wiss. Ergebn. Schwed. Zool. 
Exped. Kilimandjaro, II, pt. 10, p. 3; 1908, Denkschr. Med.- 
Naturw. Ges. Jena, XIII, pt. 1, p. 176. 
Speiser described the genus as new in two different pub- 
lications; but it should be dated from the first appearance 
in 1907, where it included only two species, Lipoptena sepia- 
cea Speiser (1905) and Echestypus parvipalpis Speiser 
(1907). One of these must be the genotype and I select 
herewith as such the older species, Lipoptena sepiacea. In 
1908, Speiser included Lipoptena sepiacea and Echestypus 
hinoculus Speiser (1908) ; but not E. parvipalpis. Aldrich 
(1923, Insecutor Inscitiae Menstr., XI, p. 77) selected “E. 
binoculatus Speiser” (misspelling of hinoculus) as geno- 
type; but, that species not being mentioned with the 1907 
description, Aldrich’s type designation is invalid. 
Echestypus agrees with Lipoptena in all essential charac- 
ters, save two, the absence of ocelli and the short or vestigial 
palpi. Neither of these characters is of particular phylo- 
genetic importance in Hippoboscidse, and I am inclined to 
regard Echestypus merely as a subgenus of Lipoptena. 
Within the limits of the latter genus, the length of the palpi 
varies considerably, being very short in some species. 
The three species which I recognize in Echestypus agree 
also in the following characters, all of which, however, are 
found in one species or other of Lipoptena. Head and 
thorax dorsally with few, spaced setae: one to six frontal 
bristles, none of them behind the eyes; one pair of vertical 
bristles ; mesonotum on each side with a conspicuous curved 
