1936] 
Crematog aster from Guatemala 
45 
female. Thus in Bruchomyrma acutidens Santschi, an 
Argentinian parasite of Pheidole nitidula strobeli Santschi, 
the female has large though veinless wings but the wings of 
the male are considerably reduced ; in the Congolese Aner- 
gatides kohli Wasmann, a parasite of Pheidole megacephala 
melancholica Santschi, the wings in the female are like those 
of Bruchomyrma, but the male is subapterous, and in the 
well-known European Aner gates atratulus, which parasi- 
tizes Tetramorium csespitum, the female has well-developed, 
veined wings, but the male is completely apterous. The 
final stage is represented by the Atitlan Crematogaster, in 
which, as I have repeatedly stated, both sexual castes are 
apterous. Nevertheless, the loss of the wings seems to have 
been so recent that it has affected neither the volume and 
structure of the thorax nor the volume of the wing-muscula- 
ture. Mating in this form must, of course, occur in the nest 
and between brothers and sisters (adelphogamy) , as in 
Anergates, but there can be no dissemination flight as in 
that species. The females, as soon as they are mature and 
have been fecunded, very probably wander away from the 
nest singly, and, in order to insure the rearing of their 
brood, enter and secure adoption in weak colonies of the host 
species. 
We have still to consider the taxonomic status of the Cre- 
matogasters discussed in the preceding paragraphs. The 
complete aptery of the sexual phases, whether we regard 
them as representing an independent species or as cospecific 
with the workers, is a character of sufficient importance to 
justify the erection of a new subgenus. Since I have inter- 
preted the sexual forms as workerless parasites I propose 
the name Apterocrema subgen. nov. for the males and 
females and describe them below as a new species, atitlan- 
ica. The workers, however, would belong to a new sub- 
species of sumichrasti, for which I propose the name maja, 
allied to Forel’s subsp. surdior and therefore belonging to the 
subgenus Orthocrema. Should the future investigator suc- 
ceed in demonstrating that the sexual castes and workers are 
cospecific, he might be willing to retain my subgeneric name, 
but maja would, of course, become a synonym of atitlanica. 
