1963] 
Carpenter — Megasecopteron 
49 
the main veins and cross veins. Most of them are short, robust spines, 
like those in certain Odonata (Fraser, 1942). In addition, hispanica 
has a fringe of short, stout setae extending along the distal and pos- 
terior margins of the wing (fig. B, plate 5). I have not previously 
observed a similar fringe in any Megasecoptera, although a weakly 
developed one does occur in some Palaeodictyoptera, as Dunbaria. 
The presence of numerous irregular cross veins, reminiscent of 
those of most Palaeodictyoptera, in a wing which is otherwise typically 
megasecopterous is indicative of the assumed relationship between these 
two orders. I believe that Anchineura brings the Palaeodictyoptera 
and Megasecoptera closer together than any previously known fossil 
has done. This is not to imply that the Anchineuridae are in the 
direct line of evolution between these orders — much more informa- 
tion about these insects, especially body structures, needs to be known 
— but Anchineura provides some evidence for a relationship which 
was previously only assumed. 
References 
Carpenter, F. M. 
1962. A Permian Megasecopteron from Texas. Psyche, 69:37-41. 
Fraser, F. C. 
1942. A note on the wing-trichiation of the Odonata. Ent. Mo. Mag., 72: 
235-236. 
Wagner, R. H. 
1958. Pecopteris pseudobucklandi Andrae and its generical affinities. 
Mededelingen Geologische Stichting (n.s.) no. 12:25-30 (Haar- 
lem). 
1955. Flora fossil y estratigrafia del carbonifero de Espana N.W. y 
Portugal N. Estudios Geologicos, Instituto de Investigaciones 
Geologicas “Lucas Malada”, 15:393-420. 
Explanation of Plate 5 
Anchineura hispanica, n.sp. A, photograph of part of the costal area of 
wing (area labeled a on plate 4). B, photograph of part of the hind margin 
of wing (area labeled b on plate 4). 
