1964] 
Cooper — Fossil Tardigrade 
45 
are joined at their bases to form an inner (cephalad) and an outer 
(caudad) compound branched claw, or “diplogriffe”, then the major 
(dorsal) branch of the inner pair is the longest of all the four 
elements, and the major (dorsal) branch of the outer pair is the 
second longest claw of each leg 1 . Finally, the minor element of the 
inner pair of claws is larger than the minor ramus of the outer pair. 
Whether joined in pairs or not, the four unequal elements are obvi- 
ously asymmetrically disposed with respect to the median plane of 
the leg. 
Type: MCZ No. 5213, Museum of Comparative Zoology, Harvard 
University. 
Type locality: One source of Legg’s collection was the drift on Amber 
Beach, a shore that rims a small bay at the base and to the south of 
Oleson Point. This is in fact the site from which most of the collections 
of chemawinite have come. The second source, which he believed to 
have discovered, was the drift and debris along a beach lying due west 
of Oleson Point, and due south of the southern limit of the Chemawin 
Indian Reservation 2 . These beaches are near to one another and to the 
entrance of the Saskatchewan River into Cedar Lake, and their 
secondary deposits are probably of a common primary origin. Regret- 
tably no record remains from which of the two beaches the piece 
containing Beorn leggi derived. 
A second tardigrade : In addition to Beorn leggi, the amber specimen 
contains a second, juvenile tardigrade (fig. ic, d; plate 6 , fig. 6 ). 
This specimen is displayed in dorsal and ventral views. Though 
adequate to establish its identity as a tardigrade, it is very poorly 
preserved, curled on itself and shriveled. There is so little detail 
that can be made out reliably, it does not merit description. But if 
there be lateral cirrus and clava as the full ventral view hints (fig. 
iD), it is a heterotardigrade, perhaps an echiniscoid as the two claws 
suggest which are visible from above on the left posterior leg. In any 
case it is evidently remote from Beorn in its affinities. 
Relationships: Though certain morphological features of cardinal 
taxonomic importance cannot be made out in the specimen as it is 
now displnved, including stylets, buccal apparatus with its possible 
1 The first three legs of the left side are displayed directly in end view in 
text fig. IB, and legs 2 and 3 are similarly viewed in the photograph of fig. 4, 
plate 6; regrettably the optical images given bv the refringent claw-rays of 
these legs are no more than bright beads of light at each focal level, and 
nothing can be made out by me of their union with the soles of the legs 
themselves. 
2 See sheet No. 63F, the The Pas Quadrangle, Department of Interior, 
Topographical Survey of Canada, 1927. 
