1964] 
Carpenter — Dictyoptilus 
ii3 
to the primitive families (e.g., Dictyoneuridae) of the order Palaeo- 
dictyoptera. A different view has been advanced by Laurentiaux 
( 1 95 7 ) , who associates it with the family Eugereonidae, which has 
included a single genus from the Permian of Germany, and which 
has become well known because of the presence of a haustellate beak. 
Eugereon was designated by Handlirsch (1906) the type-genus of a 
new order (Protohemiptera) but it has been included by most stu- 
dents of Palaeozoic insects in the Palaeodictyoptera. Lameere (1935), 
however, separated the group, which he termed the Pseudohemiptera, 
from the palaeodictyopterous families (including Dictyoptilidae) by 
superordinal lines. Laurentiaux (1953) has elevated the Palaeodic- 
tyoptera to a superorder and has recognized the Protohemiptera as an 
order within that complex. In addition to the Eugereonidae and 
Dictyoptilidae, Laurentiaux has included within the Protohemiptera 
the Protagrionidae (based on the monotypic genus Protagrion ) and 
the Calvertiellidae (based on the monotypic genus Calvertiella) . For 
reasons which will be apparent in the following discussion, I believe 
Laurentiaux is right in associating Dictyoptilus with Eugereon but I 
do not believe there is evidence to justify the inclusion of Protagrion 
and Calvertiella in the same complex. 
Text-fig. 1. Eugereon boeckingi Dohrn. A. fore wing; B, hind wing. 
Original drawings based on photograph of type sent by Dr. Paul Guthorl, 
Lettering as in Plate 13. 
A detailed study of the venation of Dictyoptilus indicates that this 
genus is actually closer to Eugereon than has been assumed even by 
Laurentiaux and, as I have mentioned above, I consider that the two 
belong to the same family. The basal part of the fore wing, for 
example, is strikingly similar to that of Eugereon (text-fig. 1) ; the 
stem of M arises precisely the same and forms a double vein with the 
stem of R, although in Eugereon the double vein is somewhat longer 
than it is in Dictyoptilus. The stem of Cu arises in the same fashion 
in both of these genera and CuA diverges anteriorly and touches the 
